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Receptor
structure specialized to detect a stimulus
Sense Organs
structure that combines nerve tissue surrounded by other tissues that enhance response to a certain type of stimulus
Transduction
conversion of one form of energy into another
What are the two stages in the sensory process?
Sensation and perception
Sensation
the process of when a sensory receptor detects stimulus
Receptor potential
small local electrical change
Perception
the conscious interpretation of stimuli
Sensory receptors transmit what four kinds of information?
Modality, location, intensity, and duration
Modality
type of stimulus or the perception it produces
What are examples of modality?
vision, hearing, taste
Location
where a stimulus is located (encoded by which nerve fibers are firing)
Receptive field
area within which a sensory neuron detects stimuli
What does the size of the receptive field determine?
Resolution
Resolution
ability to distinguish between two close-together stimuli
Intensity
strength of stimulus
What can a weak stimuli activate?
Most sensitive neurons
What can a strong stimuli activate?
Less sensitive neurons with higher thresholds
Which stimuli has the most fibers respond?
Intense stimuli
What happens to the rate of fibers firing as the stimulus intensity rises?
Fibers fire faster
Duration
how long stimulus lasts
Sensory adaption
if a stimulus is prolonged, firing of the neuron gets slower over time, we become less aware of it
Phasic receptors
rapidly adapt to a constant stimulus and turn off
Examples of phasic receptors?
smell, hair movement, and cutaneous pressure
Tonic receptors
adapt slowly, action potentials continue more steadily while stimulus is present
Examples of Tonic receptors?
Body position, muscle tension, joint motion
General (somatosensory, somesthetic) senses
widely distributed receptors in the skin, muscles, joints, tendons and viscera. Touch, pressure, stretch, heat, cold, pain and unperceived stimuli like blood pressure and composition.
Examples of General senses?
Touch, pressure, stretch, heat, cold, pain, blood pressure, blood composition
Special senses
limited to head, innervated by cranial nerves, and involve complex sense organs
Special sense examples?
vision, hearing, equilibrium, taste, smell
Gustation
sensory perception of molecules dissolved in water
Tastants
Chemicals that stimulate gustatory receptor cells
Taste buds
sensory cells clustered in 4,000
Lingual papillae
visible bumps on the tongue
How many Lingual papillae are there?
4
How many Lingual papillae have taste buds?
3
Filiform papillae
small spikes with no taste buds
What is the function of Filiform papillae?
Sense food texture
Foliate papillae
form parallel ridges along sides of posterior two-thirds of tongue
What is the function of foliate papillae?
have taste buds that mostly degenerate by age 3
Fungiform Papillae
shaped like mushrooms, at tips and sides of tongue
What does each fungiform papillae have?
three taste buds
Vallate papillae
large papillae arranged in a V at rear of tongue
What does the vallate papillae contain?
one-half of all taste buds
Taste Buds
clusters of 50-100 taste cells, supporting cells, and basal cells
Taste Cells
epithelial cells, not neurons
Taste hairs
apical microvilli, act as a receptor surface
Taste pore
opening in taste bud
Basal cells
stem cells that replace taste cells every 7 to 10 days
Supporting cells
resemble taste cells but no synaptic vesicles or sensory role
What are the five primary sensations of taste?
Salty, sweet, umami, sour, bitter
Salty
produced by metal ions (sodium, potassium)
Sweet
associated with carbohydrates and other foods of high caloric value
Umami
meaty or savory taste of amino acids in chicken or beef broth
Sour
acids such as in citrus fruits
Bitter
associated with spoiled foods and alkaloids such as nicotine, caffeine, quinine
What is taste influenced by?
Food texture, aroma, temperature, and appearance
Recent research suggests that there are two more primary tastes which are?
Oleogustus (taste of fats) and water
What is the process of taste for umami, sweet, and bitter?
Second messenger system: sugars, alkaloids, and glutamate activate G protein coupled receptors on taste cell membrane, activate second-messenger systems
What is the process of taste for salty?
Na+ goes straight into the cell, depolarizes it, and then serotonin and ATP are released
What is the process of taste for sour?
H+ enters and lowers pH of the cell causing for potassium to remain in the cell causing it to depolarize
What three cranial nerves carry taste information?
Facial nerve (anterior two-thirds of tongue), Glossopharyngeal nerve (posterior one-third of tongue), and Vagus nerve (palate, pharynx, and epiglottis)
Where do the fibers of the three cranial nerves involved with taste synapse?
Medulla oblongata
Second-order neurons from medulla oblongata relay to where?
Hypothalamus, amygdala, thalamus to primary gustatory cortex, and thalamus to orbitofrontal cortex
Olfaction
sense of smell
Odorants
airborne chemicals
Olfactory mucosa
patch of epithelium in roof of nasal cavity that houses receptor cells
Olfactory cell structure
neck and head are a modified dendrite, and head contains 10-20 immobile cilia
Olfactory hairs
immobile cillia
How many odorant receptors do humans have?
400
Each olfactory cell has receptors for how many kinds of chemical odorants?
one
1st step of Olfactory transduction
odorant binds g protein couple receptor on olfactory, activates cAMP secondary messenger system
2nd step of Olfactory transduction
Opens ion channels for Na and Ca which depolarizes membrane and triggers action potential that travels to the brain
1st step of Olfactory projection to the brain
Olfactory signals are received in olfactory bulbs
2nd step of Olfactory projection to the brain
Signals are relayed caudally in olfactory tracts
3rd step of Olfactory projection to the brain
Primary olfactory cortex in temporal lobe creates conscious perception of odor and relay signals to other brain destinations
4th step of Olfactory projection to the brain
Signals from each temporal lobe are relayed to contralateral temporal lobe, so all processing is mirrored in both cerebral hemispheres
Hearing
Response to vibrating air molecules
Equillibrium
Sense of motion, body orientation, and balance
Sound
any audible vibration of molecules
What causes the eardrum to vibrate?
air molecules hitting it
Pitch
whether a sound is high (treble) or Low (bass), determined by frequency of vibration which is measured in hertz
Loudness
perception of sound energy, intensity, or the amplitude of vibration
What is the typical range of frequency for sound for humans?
20-20,000Hz
Outer (external) ear
Funnel for conducting airborne vibrations to the eardrum
What is the outer cartilaginous part of the ear on the side of the head?
auricle (pinna)
Auditory canal (external acoustic meatus)
Slightly curved passage approximately 3cm long leading through temporal bone to eardrum
Outer end is protected by stiff what?
Guard hairs
Cerumen (earwax)
Ceremonious and sebaceous gland secretions mixed with dead skin cells
Functions of Cerumen?
Traps microbes, waterproofs canal, and keeps eardrum pliable
Middle Ear
Located in tympanic cavity in temporal bone, bounded by eardrum and inner ear
Tympanic membrane
The eardrum, a slightly conical membrane that vibrates in response to sound (highly sensitive to pain)
Tympanic cavity
air-filled, 2 to 3 mm wide space between outer and inner ears
Mastoid cells
air filled spaces in mastoid process of temporal bone
Auditory (eustachian or pharyngotympanic) tube
passageway to nasopharynx, normally flattened and closed but opens when swallowing or yawning
Auditory ossicles
three smallest bones of body, connect tympanic membrane to inner ear
Malleus
has long handle attached to inner surface of tympanic membrane and a head that articulates with incus
Incus
has triangular body that articulates with malleus and long limb articulates with stapes
Stapes
shaped like a stirrup; footplate rests on oval window
Oval window
where inner ear begins
Stapedius muscle attaches to what?
Stapes