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Biodiversity
the total variety of organisms, habitats, communities and ecological processes in the biosphere
High biodiversity
the ecosystem is healthy and more likely to withstand sudden changes
Low biodiversity
the ecosystem is susceptible to small change and likely to collapse
3 factors making up biodiversity
genetic, species, ecosystem
Species richness
number of different species represented (count of species)
Species richness index
D = S/√N (no. species / √no. individuals)
Evenness (relative abundance)
a description of the number of individuals of a species in relation to the total number of individuals in an area
Percentage cover
proportion of an area covered by an organism (i.e.: estimating plant frequencies)
Advantages of percentage cover
good estimate of abundance, more efficient than counting individuals
Disadvantages of percentage cover
plants in flower overestimated, low-growing plants underestimated, poor quadrat sampling means some rare species can be missed
Emergent layer
tallest / most predominant layer, produces the most above-ground biomass
Percentage frequency
appearance of plant species in sample quadrats
% freq = [no. quadrats containing species]/[total no. quadrats]
Simpsons Diversity Index
measure of diversity that takes into account the number of species and relative abundance
D = 1 - ([Σn(n-1)]/[ΣN(N-1)])
Intraspecific interactions
interactions between the same species
Interspecific interactions
interactions between different species
Predation
predator kills prey for food
Competition
organisms competing with one another for limited resources (food, space, mates)
Symbiosis
2 different species living together in direct contact (3 forms)
Mutualism (symb.)
both species benefit (++)
Commensalism (symb.)
one species benefits, other unaffected (+0)
Parasitism (symb.)
one species benefits, other harmed (+-)
Disease
any condition that impairs the normal activity of an organism (infectious & non-infectious)
Abiotic factors (examples)
Climate - rainfall, temperature, wind, sunlight
Substrate
Soil - structure, porosity, pH, salinity
Size / Depth of area
Biotic factors (examples)
Types of plants & animals
Inter/intraspecific species interactions
Spatial scale
space, geographical location... spatial patterns of distribution
Temporal scale
time... change occurring over a period of time
Threats to the Great Barrier Reef
Climate change - temperature, rising sea levels
Land based run-off - pesticides, detergents
Coastal development - tourism, urban development
Illegal fishing - interrupting food chains
Distribution
region where an organism is found
Abundance
density or no. individuals in that area
Factors affecting distribution & abundance
Physical - temperature, rainfall, altitude, light, soil
Biotic - food source, predators, competitors, etc.
Underpopulation - some species need min. pop. size
Chemical - pH, salinity, water
Dynamic - wind speed/patterns, wave actions
Taxonomy
the science of classifying organisms based on shared characteristics
Binomial classification
systematic classification and description of species by 2 Latin names (using binomial nomenclature)
Benefits of a classification system
Understand relationships, evolution & diversity
Internationally recognised (easy communication & access to quick information)
Linnaean system - classification system
classification based on similarity and functions of physical features
4 features of the Linnaean System
Hierarchical structure
3 kingdoms - Eukarya, Archaea, Eubacteria
Uses binomial nomenclature
Based on morphological features
Advantage of the Linnaean System
can distinguish major groups
Limitation of the Linnaean System
no information about evolutionary relationships
Linnaean System - Taxa for Eukaryotes
Kingdom
Phyla (inv. PCPAMA) (vert. BFARM)
Class
Order
Family
Genus
Species
Methods of Reproduction - classification system
classification based on how species reproduce
Sexual reproduction (methods of reproduction)
union of 2 gametes from 2 parents
External fertilisation - outside female
Internal fertilisation - inside female
Hermaphrodites - male + female organs in 1 body
Asexual reproduction (methods of reproduction)
one parent producing identical offspring
Parthenogenesis (without fertilisation by sperm)
Survivorship curves
graphical representation of species survival patterns as a function of age
Type I (survivorship curve)
high - low survivorship... few offspring receive care and nurturing (i.e.: large mammals)
Type II (survivorship curve)
steady survivorship... constant death rate over lifespan (i.e.: small rodents)
Type III (survivorship curve)
low - high survivorship... many offspring that receive little care (i.e.: fish)
Reproductive strategies (r/K selection theory)
decide the quality & quantity of offspring produced by a species
K strategy (r/K selection theory)
type I survivorship (high-low)... predictable environment, small numbers of offspring
r strategy (r/K selection theory)
type III survivorship (low-high)... unstable environment, large numbers of offspring
Molecular Sequences - classification system
systemics / molecular phylogeny - using genetic information (i.e.: DNA sequences) and molecular differences to determine relationships between organisms, both present and extinct
Phylogenetic tree
shows evolutionary relationships between both present and extinct organisms
branch lengths proportional to amount of inferred evolutionary change
CLADE
A GROUP OF ORGANISMS THAT CONSIST OF A COMMON ANCESTOR AND ALL ITS LINEAL DESCENDANTS
Cladogram
a diagram that shows evolutionary relationships between species
Assumptions of Cladistics
1. Common ancestry - all species diverged from the same common ancestor
2. Bifurcation - existing species evolved into 2 groups (2 new lineages)
3. Physical change - characteristics of organisms change over time
Biological Species Concept (BSC) - definition of species
a species is a group of organisms that can interbreed and produce fertile offspring
Limitations of the BSC
Can't test production of fertile offspring with fossils
Species don't always sexually reproduce
Some hybrids can be fertile
There is a flow of genetic material that does not involve (sexual/asexual) reproduction
Phylogenetic Species Concept (PSC) - definition of species
a species is the smallest set of organisms that share a common ancestor and can be distinguished from other sets
Limitations of the PSC
Can't apply the PSC to all forms of life
Very few groups of organisms have detailed evolutionary histories
Mules (Equus mulus) - female horse x male donkey
an interspecific hybrid that does not produce fertile offspring
Microhabitat
a small, specialised habitat within a larger habitat with unique conditions (sometimes home to unique species)
Ecosystems composed of varied habitats
microhabitat - habitat - ecosystem - biome
Ecosystem classification - management
Ecosystem classification is important in devising policies & strategies for effective ecosystem management to provide ecosystems with long-term resilience
Importance of old growth forests
provide a high level of biodiversity, maintain a wide range of ecological functions
Old growth forest management strategies
prescribed burning, government protection from logging
Importance of soil productivity
produces critical and essential macronutrients, very important for effective ecosystem management
Soil productivity management strategies
monitoring conditions
Importance of coral reefs (i.e.: GBR)
provide a megadiverse marine environment with high biodiversity
Coral reef management strategies
prohibit / limit the fishing of certain species, reduce land based run off and coastal development
Stratified sampling
dividing a population into non-overlapping subgroups (strata), then taking random samples from each stratum
Benefits of stratified sampling
- reduced error of estimation
- increased prevision for same sample size
- reduced cost of sampling
Disadvantages of stratified sampling
- population must be divided into clearly identifiable strata
- no accurate estimation of proportions of each stratum
- some strata must be joined to ensure adequate numbers
Random sampling
samples / quadrats randomly selected to record data on a population as a whole
Random vs Stratified sampling
Random sampling selects areas or parts of a population randomly without any other consideration to collect data that represents an entire population, whereas stratified sampling first divides the population into subgroups (strata) that are homogenous within, but vary between stratum, before recording data. Stratified sampling ensures that all layers or stratum are included in data analysis, whereas random does not.
Transect
a line/strip/profile for counting & mapping the number of individuals at different distances (line transect, belt transect, profile diagram)
Purpose of a transect
to identify the effect of an abiotic factor on the abundance or distribution of organisms
Quadrat
a square grid used to estimate the abundance of slow-moving or plant species in an area
Purpose of a quadrat
to determine the distribution, abundance & density of a population when total counts can't be made
4 steps to minimising bias
1. law of large numbers (more samples = increased reliability of data)
2. random-number generator
3. counting criteria (a consistent counting method
4. calibration of equipment
Autotrophs (producers)
produce all organic compounds in ecosystems by photo/chemosynthesis
Heterotrophs (consumers/decomposers)
obtain organic compounds by consuming autotrophs (primary) or other heterotrophs (secondary)
Carbon fixation
the fixation of inorganic carbon into organic molecules by autotrophs
The transfer of chemical energy along a food chain
10% transferred to next trophic level
90% used to power organisms metabolic processes
Primary productivity
productivity of autotrophs (primary producers)
Secondary productivity
production of organic matter by heterotrophs due to assimilation of food and synthesis of new cells/tissues
Gross productivity
total amount of organic matter produced
Gross primary production (GPP)
total amount of CO2 fixed by plants in photosynthesis
Net primary productivity (NPP)
the amount of energy that plants pass on to the next trophic level (after respiration and other metabolic processes have occurred)
NPP = GPP - R
Net ecosystem productivity (NEP)
amount of energy remaining after the costs of respiration by autotrophs and heterotrophs
NEP = GPP - (R0 + R1 + R2...)
Water cycle
transpiration/runoff - evaporation - condensation - precipitation - infiltration
Carbon cycle
atmosphere (respiration/combustion) - absorption (by autotrophs) - assimilation (eating along food chain) - decomposed (decomposers) / combusted (fossil fuels)
Nitrogen cycle
1. fixation (nitrogen fixing bacteria; nitrogen - ammonia/ammonium... nitrifying bacteria; ammonium - nitrate/nitrite... OR lightning)
2a. denitrification (denitrifying bacteria; nitrates/nitrites - atmospheric nitrogen)
2b. absorption (plants absorb nitrates/nitrites)
3. assimilation (nitrogen along food chain)
4. ammonification (decomposers; return nitrogen to soil in ammonia
ECOLOGICAL NICHE
THE ROLE AND SPACE THAT AN ORGANISM FILLS IN AN ECOSYSTEM, INCLUDING ALL ITS INTERACTIONS WITH THE BIOTIC AND ABIOTIC FACTORS OF ITS ENVIRONMENT
Factors determining the ecological niche of an organism
Habitat - the space an organism lives
Feeding relationships - food and food avaliability
Interspecific interactions
Competitive Exclusion Principle
2 species cannot occupy the same ecological niche (competing for the same limited resources)
If 2 species occupy the same niche, competition will show that one species has a competitive advantage. The less-favoured species must either adapt (natural selection) or will become extinct
KEYSTONE SPECIES
A PLANT OR ANIMAL THAT PLAYS A UNIQUE AND CRUCIAL ROLE IN THE WAY AN ECOSYSTEM FUNCTIONS
Importance of keystone species
- occupies a crucial role in the food web of an ecosystem
- removal of the keystone species would have a dramatic effect (proportions of species would be imbalanced)
- removal of the keystone species will likely cause the ecosystem to collapse
Examples of keystone species
Cassowary - eats fruit and is main dispersal agent for many plants
Northern Quoll - keystone predator (eats insects, reptiles, small mammals, and fruits)
Acorn Banksia - only source of nectar for honeyeaters at some times of year (honeyeaters important for pollination of plants)
CARRYING CAPACITY
THE SIZE OF THE POPULATION THAT CAN BE SUPPORTED INDEFINITELY ON THE AVAILABLE RESOURCES AND SERVICES OF AN ECOSYSTEM
Biotic factors limiting carrying capacity
- competition for resources
- predation
- disease
Abiotic factors limiting carrying capacity
- space
- availability of nutrients
- pollution / natural disasters
- extreme climactic events (i.e.: drought / temp. change)
Density dependent factors limiting carrying capacity (determined by population size)
- competition for resources
- predation
- space / crowding
- infectious disease / parasitism