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Type of plant defenses
Physical barriers: Waxes, bark,cork
Chemical Defenses: 3 major groups
Pathogen-specific compounds: SAR and phytoalexins
1st tier
waxes
Cuticle of the epidermis
waxy layers in the root: endodermis
bark is heavily waxy to prevent bacteria and bugs from attacking the plant
2nd tier
chemical compounds
defense cpds
3 groups: N2 containing cpds, terpenoids, phenolics
N2 containing compounds
alkaloids, cyanogenic glycosides, glucosinolates expensive to make( in terms of ATP and N, remember N is hard for plants to get)
Alkaloids
>6000 known alkaloids
about 20% of flowering plants make these anti insect compounds
nicotine
caffeine
codeine
cocaine
capsaicin
Cyanogenic Glycosides
25 known
>2600 plant species know to produce cyainde gas
components kept separate until herbivore damage occurs
cassava
crabapples
cherries
apples
glucosinolates
cabbage family (broccoli,cabbage)
composed of 2 parts a and b
terpenoids
made of 5 paired carbon units called isoprenes
menthol
pyrethroids
gossypol
made from acetyl-coa or compunds found in respiration pathways
are the largest group of secondary metabolites
GA is a diterpene, carotenoids are tetraepenes, and ABA id sesquiterpene
are toxins and feeding deterrents
phenolics
made of long chain phenol groups
make plants bitter tasting
aspirin
tannin
lignin
3rd tier
specific responses
Hypersensitive response: dead tissue response
pathogenesis-related genes; activated when pathogen proteins bind to receptor protein and activate genes for systemic acquired resistance (pheromone stimulates salicylic acid production in other plants), phytoalexins ( antimicrobial compounds; cant be detected until infection occurs)
Plant stress physiology
stress reduces crop yields by up to 75%
major stressors: heat, cold, water, salt
heat stress
destablizes lipids of phospholipid bilayer, allowing permeability to increase dangerously enzymes shut down
heat shock protein (HSP) stabilizes these enzymes
how do plants get rid of potential heat load
for sun leaves heat load is very high
it has been calculated that a 0.3mm thick leaf would heat up to 100 degree C every minute if the light absorbed was not converted into sugars and heat
ways to get rid of heat from plants
re emission as red light
evaporative heat loss
by isoprene production (stabilize photosynthetic membranes under hotter conditions inside the leaf)
xanthophylls (yellow accessory pigments) can cycle by conversion from one to another, resulting in a dissipation of heat rotate their chlorplasts up and down to longer palisade cell
cold stress
Ice crystal formation causes damage; crystal formation inside the cell is less damaging than between cells
At cold temps, membranes solidify. Some plants produce anti-freeze compounds that keep membranes from solidifying
water stress effects
leaf expansion
turgor decreases
cell wall chemistry/ structure changes
leaf abscission
increased root expansion
stomatal closure due to ABA production ( thus carbon intake stops)
salt stress
salts can change water potential so that water moves out of plant to soil
halophytes pull salt ions out of incoming water before that can affect the plants biochemistry
weather
short term
snapshot of current conditions
climate
long term
decades/centuries of measured weather conditions
climate components
atmosphere: air flow, 78% nitrogen, 21% oxygen
Hydrogen: water flow/presence
Geosphere: Earth crust/volcanoes
Biosphere: influence of the biological processes
Cryosphere: influence of the ice/snow/glaciers
Sources of ancient climate data
tree rings can gice us O16/O18 ratios, C12/C13/C14 ratios
Ice cores from glaciers
coral reef cores
cave deposits
effects of a warming earth on crops and livestock
Nitrogen and carbon cycles are altered by increased CO2
Increased nitrogen efficiency use by crops by 19%
biological nitrogen fixation increase of 55%
lower nitrogen fertilizer requirements
higher CO2 levels save water in crop plants
Higher CO2 levels are predicted to: increase the efficiency of carbon uptake by crop plants, save water as plant gets carbon, doesn’t have to leave stomata open as long
Effects on pathogens
All pathogen groups are predicted to increase substantially, both in numbers and in range locations
increase in the number of pathogens and new ranges will offset any increase in crop yields