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19.1
overview of evolution
evolution
a change in one or more heritable characteristics of a population from one generation to the next
natural selection
environment
artificial selection
human hand
Darwin
influenced by work in other fields and his own observations on the Beagle (his boat) during about 5-year journey
Darwin noted distinctive traits of island species that
allowed them to better exploit their environment
darwin suggets
that existing species are derived from pre-existing species
Darwin expressed his ideas about evolution as
“the theory of descent with modification through variation and natural selection “
variation
traits heritable - passed from parent to offspring
genetic basis was not yet known in Darwin’s time
natural selection
more offspring are produced than can survive
competition for limited resources
Individuals with beneficial heritable traits are more likely to survive and reproduce
as a result of natural selection,
certain traits that facor reproductive success become more prevalent in a population over time
19.2
evidence of evolutionary change
selective breeding
the english bulldog
dogs
convergent evolution
similar DNA because of a similar habitat
2 species from different lineages have independently evolved similar
analogous structures
Fossils show
evolutionary change
transitional form
biogeography
The study of the geographic distribution of extinct and living species
homology
refers to a similarity that occurs due to descent from a common ancestor
may involve anatomical, developmental, or molecular features
vestigial structures
anatomical features that have no current function but resemble structures of presumed ancestors
developmental homology
refers to similarities that occur during development
molecular homology
refers to similarities that occur at the molecular level
The degree of similarity between genetic sequences from different species reflects the evolutionary relatedness of those species
19.3
genes in populations
gene pool
includes all of the alleles for every gene in a population
polymorphism
refers to the presence of two or more alleles at a gene locus for a given character within a population
ex. two color variations
the DNA difference between alleles
large healthy populations exhibit
high levels of genetic diversity
every animal, human have __ alleles out of __
2
4
how can genes become polymorphic?
deletion, duplication, or change in a single nucleotide
single-nucleotide polymorphism SNP
The smallest type of genetic variation that can occur within a gene and is the most common
population
a group of individuals of the same species that occupy the same environment at the same time and can interbreed with one another and produce viable offspring
genetic variation
can be analyzed quantitatively
Calculations of allele frequency and genotype frequency are fundamental when analyzing populations
allele frequency =
number of copies of a specific allele in a population / total number of all alleles for that gene in the population
genotype frequency =
number of individuals with a particular genotype in a population / total number of imdividuals in the population
genotype frequency
calculated from observational data of phenotypes in a population
allele frequencies
calculated from known genotype data
hardy-weinberg equation
describes the relationship between allele and genotype frequencies when a population is not evolving
fundamental principle in population genetics
allows pop gens to determine the baseline allele frequencies in a population
hardy-weinberg law
calculate the dominant allele frequency
slides 33-34
hardy-weinberg equation describes the relationship between
allele and genotype frequencies when a population is not evolving
p + q = 1
and
p² + 2pq + q² = 1
if they do, the population is not evolving → stable
if they don’t, pop is evolving, figure out why?
equilibrium =
population NOT evolving
no new mutations
populations is so large allele frequencies do not change
no migration
no random mating
in reality, populations rarely reach equilibrium
if frequencies are not in equilibrium, an evolutionary mechanism is a work
microevolution
used to describe changes in a population‘s gene pool
(such as changes in allele frequencies)
from generation to generation
genetic drift
random chance
“accident“
19.4
natural selection
over time, natural selection may result in
adaptations
changes in populations of living organisms that increase their ability to survive and reproduce in a particular environment
as a result of natural selection
certain traits that favor reproductive success become more prevalent in a population over time
reproductive success
the likelihood of an individual contributing fertile offspring to the next generation
fitness (w)
The relative likelihood that one genotype will contribute to the gene pool of the next generation compared with other genotypes
Fitness is a measure of reproductive success
directional selection
individuals at one extreme of a phenotypic range have a greater reproductive success in a particular enviornment
stabilizing selection
Favors the survival of individuals with intermediate phenotypes and selects against those with extreme phenotypes
diversifying (disruptive) selection
Favors the survival of two or more different genotypes that produce different phenotypes
balancing celection
maintains genetic diversity in a population
sexual selection
a form of natural selection by which individuals with certain traits are more likely than others to engage in successful mating
intrAsexual selection
members of one sex (usually males) directly compete with each other for the opportunity to mate with individuals of the opposite sex
intErsexual selection
members of one sex (usually females) choose their mates on the basis of certain desirable characteristics
19.5
genetic drift
genetic drift
refers to the changes in allele frequencies due to random chance
over many generations, genetic drift favors either
elimination or the fixation of an allele
genetic drift is
buffered in large populations
But it can create a huge impact on smaller populations
bottleneck effect
refers tio the change in allele frequencies of the resulting population due to genetic drift
founder effect
occurs when a small group of individuals separates from a larger population and establishes a new colony in a new location
gene flow and migration
enhance genetic diversity