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adventitious roots
Refers to an organ that forms in an unusual place; refers primarily to roots that form on leaves, nodes, or cuttings rather than on another root
buttress roots
A root that grows asymmetrically such that it becomes very tall and thin, extending up the trunk as much as a meter or more and giving the trunk lateral stability
casparian strips
A layer of impermeable lignin and suberin in the walls of endodermal cells, preventing diffusion of material through that portion of the wall
ectomycorrhizal relationship
A type in which the fungi invade only the outermost cells of the root
endodermis
A sheath of cells surrounding the vascular tissue of roots (and occasionally horizontal stems); their Casparian strips prevent uncontrolled diffusion between root cortex and root vascular tissue by means of walls and intercellular spaces
endomycorrhizal association
A type in which the fungi invade all cells of the root cortex
fibrous root system
A set of roots in which all roots are more or less the same size; that is, none is especially larger than any other root (none is a taproot)
haustoria
The structure by which a parasite enters and draws nutrients from a plant; in fungi, it is a hypha; in mistletoes and similar parasites, it is a modified root
lateral roots/branch roots
Any root that is borne by another root (i.e., any root that is not attached to a stem or other nonroot organ)
order of bark growth
cork - cork cambium - 2 phloem - vascular cambium - 2 xylem
mycorrhizae
Fungi that form a symbiotic relationship with roots, usually of benefit to plants because they provide phosphorus
what does the periderm consist of?
cork cambium + layers of cork + any cells to the inside from cork cambium
nitrogen fixation
The conversion of atmospheric nitrogen into any compound that can be used by plants, typically either nitrate or ammonium
pericycle
An irregular band of cells in the root, located between the endodermis and the vascular tissue
prop roots
Any root that grows through air or water before entering soil; prop roots help to brace and stabilize a shoot
quiescent center
Portion of the root apical meristem in which cell division does not occur
radicle
The main root of a seed; it is the direct continuation of the embryonic stem
root apical meristem
The set of cells capable of dividing, located at the apex of a root. Produces the root itself and also the root cap
Root cap
A layer of parenchyma cells that cover and protect the root apex.
root hairs
A short-lived trichome near a root apex. Root hairs increase the surface area and absorptive capacity of roots; being single, slender cells, they enter soil pores that are too small for whole root tips to enter
root nodule
Several species of plants have symbiotic relationships with nitrogen-fixing bacteria; in some species, especially those of legumes, portions of roots become swollen and occupied by the bacteria
taproots
Technically, a taproot is only the root that develops from the radicle (the embryonic root). Very commonly the term “taproot” is used to refer to any root that is markedly larger than most other roots
zone of elongation
Region of a root tip, just proximal to the root apical meristem, where cells undergo pronounced elongation
wood
secondary xylem
vascular cambium
The meristem that produces secondary vascular tissues—secondary xylem (wood) and secondary phloem (inner bark)
tylosis
After a vessel stops conducting because of cavitation, adjacent cells may push cytoplasm into the vessel through pits, plugging the vessel
storied cambium
A vascular cambium in which fusiform initials are aligned horizontally. Alternative: nonstoried cambium
softwoods
A term applied to both gymnosperms and their wood, because few gymnosperms have any fibers in their wood
secondary xylem
Xylem derived from the vascular cambium
secondary vascular bundles
In monocots with anomalous secondary growth (such as Joshua trees), secondary vascular bundles are those produced by the cambial zone.
secondary tissues
The tissues of the secondary plant body-those produced by the vascular cambium and the cork cambium. Alternative: primary tissues.
secondary phloem
Phloem derived from the vascular cambium
sapwood
The light-colored, light-scented outermost wood of a trunk or branch; conduction is still occurring and many wood parenchyma cells are alive. Alternative: heartwood.
ring porous
Wood in which the early wood has more numerous and larger vessels than the late wood. Alternative: diffuse porous wood
reaction wood
Wood formed in response to mechanical stress. tension wood The reaction wood of eudicots, formed on the upper side of a branch. compression wood The reaction wood of gymnosperms, formed on the lower side of a branch.
rays
In secondary xylem and phloem, a radial series of cells produced by ray initials. Alternative: axial tissue
ray tracheids
Horizontal tracheids in the secondary xylem rays of gymnosperms
ray initials
In a vascular cambium, the short cells that give rise to the rays of the secondary xylem and phloem. Alternative: fusiform initials
radial system
The set of rays within wood
primary tissues
The tissues derived more or less directly from an apical meristem or leaf primordium; the tissues of the primary plant body. Alternative: secondary tissues
phellogen
Synonym for cork cambium
phellem cell
Technical term for cork
periderm
Technical term for bark; it consists of cork, cork cambium, and any enclosed tissues such as secondary phloem
perclinal wall
A wall that is parallel to a nearby surface, especially the outer surface of the plant. Alternative: anticlinal wall
nonstoried cambium
A cambium in which the fusiform initials are not aligned horizontally. Alternative: storied cambium.
lenticels
In bark, a region of cork cells with intercellular spaces, permitting diffusion of oxygen into inner tissues.
late wood
Synonym for summer wood.
included phloem
In certain types of anomalous secondary growth, patches of secondary phloem may be located within the secondary xylem
heartwood
The colored, aromatic wood in the center of a trunk or branch; all the wood parenchyma cells have died and no water conduction is occurring. Alternative: sapwood
hardwoods
The colored, aromatic wood in the center of a trunk or branch; all the wood parenchyma cells have died and no water conduction is occurring. Alternative: sapwood
growth ring
Synonym for annual ring in wood. “Growth ring” is preferred because rings are occasionally not strictly annual (sometimes two are produced in 1 year, and none might be produced in adverse years).
fusiform initials
In a vascular cambium, the long cells with tapered ends that give rise to axial cells of the secondary xylem and secondary phloem. Alternative: ray initials.
establishment growth
In palms, seedlings initially grow wider while remaining short; during this establishment growth, the seedling establishes additional adventitious roots.
early wood
synonym for spring wood.
diffuse porous
Wood in which the vessels of late wood are about as numerous and as wide as those of early wood. Alternative: ring porous wood.
cork cell
A cell in bark that has walls encrusted with suberin; cork prevents loss of water through the bark and prevents entry of pathogens.
cork cambium
A layer of cells that produces the cork cells of bark. Also called phellogen
bark
In some plants, especially woody, perennial plants, the epidermis is replaced by bark, which always contains at least cork cells produced by cork cambia; bark may also contain secondary phloem, sclereids, and fibers (bast fibers).
axial system
In secondary xylem (wood) and secondary phloem, the axial system consists of the cells derived from fusiform initials; most cells in the axial system (e.g., tracheids, vessel elements, fibers, sieve tube members) are elongate parallel to the long axis of the wood or phloem. Alternative: ray system.
anticlinal walls
A wall perpendicular to a nearby surface, especially the outer surface of the plant. Alternative: periclinal wall.
annual ring
In secondary xylem, the set of wood, usually early wood and late wood, produced in 1 year.