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Individual
individual traits determine response to environmental factors
Population
-interacting group of conspecific individuals
-group of individuals of sam type of organisms that interbreed
Species
groups of the same type of organisms and descend from common ancestor
Community
set of co-occurring interacting species
Ecosystem
interacting species and abiotic environment
Evolution
change in genetic composition of population over time
Natural selection
differential survival and/or reproduction of individuals with different trait values
Adaptation
-structural/ physiological/ behavioral traits that enhance organism's chances of survival and reproduction in its environment
-trait evolved to enhance organism's survival and reproduction in an environment
-evolutionary change in genotype that increases performance (i.e. process by natural selection)
Estimate total number N for species
N/K=n/k
Species richness
number of species in given area
Species evenness
degree that species are equally abundant
True Diversity equation
D=(p1^-p1)(p2^-p2). . . . (pn^-pn)
Climate
measure of average pattern of variation in temperature, precipitation, and other meteorological variables in given region over long periods of time
Biomes
world's major communities, classified according to predominant vegetation and characterized by adaptations of
Chapparal
temperate shrub land/woodland with:
-hot/dry summer
-wet/cool/moist winter
-dense vegetation and vulnerable to summer fire
Ferrel cell
driven by 60 degree N (clockwise) wind current
Hadley cell
driven by 30 degree N (counterclockwise) wind current
-cool/dry air descending
Coriolis effect
-pushes floating object away from equator and will appear, relative to the earth, to pick up speed and moves east
-push floating object toward equator and will appear, relative to the earth, to lose speed and moves west
-I.E. deflection of air/water as result of differences in Earth's rotational speed at different latitudes
Rain shadow
-North/ South mountain ranges, so precipitation has rising air cooling/condensing while dry air cools/sinks (to desert in valley)
Intertropical Convergence Zone (ICZ)
-two Hadley cells working
-moves North in summer
-moves South in winter
Endemic
occurs in one particular location and nowhere else on Earth
Dispersal patterns
1. regular (uniform) dispersion
2. random dispersion
3.clumped dispersion
Motile
moves from place to place
i.e. animals
Sessile
stays in one place
i.e. still animals
Mycorrhizae
association of fungi with plant roots (>90% terrestrial plants)
-mutualism between plant and mycorrhizae (mycorrhizae not good when high water/nutrients => becomes parasitic)
Cost-benefit Approach
assumes limited time/ energy and used to investigate relationships between behavior, environment, and fitness
Principle of Allocation
all life functions cannot be simultaneously maximized
Trade-off
relationship between benefits of a trait in one context and its costs in another context
Gross Photosynthesis
-changes in O2 in light= photosynthesis-respiration
-changes in O2 in dark= -respiration
-(light change in O2)-(dark change in O2)
Liebig's Law of the Minimum
production only occurs at rate permitted by most limiting factor e.g. sunlight or water or nutrients
Root to Shoot ratio
ratio of below ground to above ground biomass
-Tundra has largest root to shoot ratio
-Tropical forest has lowest root to shoot ratio
Acclimation
change in phenotype within individual's lifetime to increase performance (is often reversible)
C4 and C3 plants
-C4 plants more specialized, but transport of C4 acid to bundle sheath cells is costly
-C3 benefiting from climate change (increases of CO2 and temperature)
Fundamental niche
set of environment conditions that individuals of a species can grow and produce
Strength/ Size Proportions
M^1/3 aproximately = L
Danger index (for surface area/size)
1/L
Metabolic rate
amount of energy expended daily at rest
-metabolic rate/ body mass increase with each other (slope =3/4 for these i.e. 3/4 power rule)
2/3rd law
strength scaling with size
Isometric scaling
all dimensions increase the same amount as the size of the organism changes
Allometric scaling
disproportionate growth of a part or parts of an organism as the size of the organism changes
Density
number of individuals in an area of quadrat
-equation is N/M= n/m
M-captured
N-estimated population size
n-recaptured
m-marked
Population Density and Average Body Size
-decrease
-slope is -3/4
Per-Capita Growth rate
r= b-d
Exponential Growth
N(t)= N0e^rt
-continues to increase
Logistic Growth
-populations may initially grow rapidly when the populations small
-then density-dependent birth/death rates cause r to decline towards 0
-pattern of population growth
Density-dependent
Ex. fire, fecundity, mortality, growth, etc.
-factors include limiting resources, predators, and pathogens
Density-independent
Ex. hurricanes, flash floods, etc.
Allee effect
some populations grow better, with some limit, when population densities are higher rather than lower
Type II Survivorship Curves
probability of surviving to next year is independent of age
-life expectancy is independent of age
Type III Survivorship Curves
high juvenile mortality but low adult mortality
-life expectancy gets better with age
Type I Survivorship Curves
low juvenile mortality but high adult mortality
-life expectancy get worse with age
Life Expectancy at Birth
lx all added together i.e. (l1+l2+l3+. . .+lx)
Life Expectancy at a Certain Age
(l1+l2+l3+. . .+lx)/ lx (lx with x being the age Ex. age 3 = l3)
R0
average number of offspring produced by individual in its lifetime
R0= l1m1+l2m2+l3m3+. . .+lxmx
R0 >1 population increasing
R0 <1 population decreasing
Developing Countries Demographic Transition
stage 2- rapid growth
stage 3- slow growth
stage 4- zero growth
stage 5(?)- negative growth
Toy Model
-population of N individuals where each individual produces 2 offspring then dies
-Each offspring survives to next year with probability of 0.5
Delayed Density-dependence
per capita growth rates depend on past densities
Vestigial structures
Ex. hind limbs in whales OR tailbones in humans
Darwin inferences
1. struggle for existence
2. natural selection
3. adaptation (evolved by natural selection)
evolutionary response (R)
-change in trait mean across generations
R= Trait mean(offspring)- Trait mean (parents prior to selection)
R=h^2S
h^2- heritability
S- selection differential
1. find S or/and h^2
2. plug into equation
3.get R; R never > S and h^2 never >1 or <0 (h^2 as 0 means no R)
Phenotype plasticity
expression of different phenotypes by the same genotype in different environments
-specific Ex. acclimation (improves performance in an environment)
Directional Selection
favors individuals with high/low trait values
-causes change in population trait mean (usually reduces variance)
-acts on individual phenotypes to change population trait mean (and frequency of alleles controlling that trait)
Stabilizing Selection
favors intermediate trait values, decreasing variance
-reduces variance but doesn't change trait mean
Ex. human birth rate
Disruptive Selection
favors extreme trait values, increasing variance
-can lead to bimodal distribution
Sexual selection
selection for traits that enhance reproductive success
-competition among members of same sex
-mate choice by opposite sex
altruist
increase fitness of other individuals at expense of its own fitness
Coefficient of Relationship
(benefit to relatives) x (coefficient of relationship) > (cost to performer)
Genetic Drift
random changes in allele frequencies from one generation to the next generation
-may produce large change in allele frequencies over time
-increasing drift in small populations and can cause loss of allelic diversity and evolutionary change as population size decreases
Population Bottleneck
situation where normally large populations may pass through environmental events that only a small number of individuals survive
Founder effects
resulting change in genetic variation
-small population size unlikely to possess all alleles found in gene pool of its source population
Semelparity
only reproduces once in lifetime
Iteroparity
reproduces more than once in lifetime
r-strategists
species with life history strategies allowing for high population growth rates
-life is uncertain
K-strategists
species with life history strategies allowing them to persist at or near carrying capacity (K) of their environment
-adapted to predicted conditions
Law of Segregation
-when individual produces gametes
-then two copies of the gene segregate and gametes only receive one copy
-Punett Square
Locus (loci)
physical location of gene on chromosome
Multiplication Rule
p1(event 1) x p2(event 2)
Addition Rule
probability of event occurring in two different ways is the sum of individual probabilities
Law of Independent Assortment
-alleles of different genes assort independently during gamete formation
-dihybrid cross (#/16)
Closeness of Loci on Chromosome
-parental gametes more frequent
-probability of recombination of loci that are closer is less likely
3 loci ====> 1. find parental gametes (most common)
2. find double crossovers (least common)
3. allele in middle "switched"
Pleiotropy
allelic variation at one locus affects multiple traits
Ex. pigmentation/hearing in cats
Antagonistic Pleiotropy
occurs when one allele has both positive and negative effects
Ex. Margan's syndrome OR cystic fibrosis
-fitness trade-off across environment
Epistasis
-phenotypic effect of allele at one locus depends upon genotype of allele at another locus
-two genes interacting in developmental or biosynthetic pathway
Ex. labardor fur OR chicken combs
Genotype x Environment Interaction
-phenotypic effect of allele or genotype depends on environment (fitness trade-off)
-geological variation in selection can maintain variation in a species
Genotypic Frequencies
f(rr)= q^2
f(RR)= p^2
f(Rr)= 2pq
Allelic Frequencies
p= f(RR)+1/2f(Rr)
q=f(rr)+1/2f(Rr)
p+q=1 OR 1-p=q ***always adds to 1!***
Prout Square
use allelic frequencies to get genotypic frequencies for a population
Hardy-Weinberg Equilibrium Conditions
1. no new mutations
2. large (infinite) population
3. random mating
4. no gene flow from other population
5. no natural selection
Disassortative Mating
between dissimilar genotypes
heterozygote excessive in newborns
Inbreeding Depression
reduced biological fitness arising from mating close relatives that tend to have the same recessive, sometimes deleterious alleles
Gene Flow
migration of individuals and/or gametes (via propagules such as seeds, etc.)
Recombinant Frequencies
proportions calculated by dividing the number of recombinant progeny by total number of offspring
Heterozygote Advantage
H-W in newborns; increasing heterozygotes with age
heterozygotes have higher fitness
-disease resistance
-antagonistic pleiotropy
-metabolic pathways
Disease Resistant (and polymorphisms)
different resistance alleles may confer resistance against different pathogens
-cystic fibrosis/cholera toxin
-Tay-Sachs disease/ TB
-Phenylketonuria/decreased chance of miscarriage from fungal toxin
Ammensalism
one species harmed and one species unaffected (prey vs. predator)
Antagonistic Interactions
one species benefits and other species is harmed
Mutualism
both species benefit from interaction
Commensalism
one species benefits and other is unaffected
Intraspecific Competition
competition between same species members
-self-thinning in plants
-Scaling law M= N^-3/2 (N decrease/M increase M =>biomass)