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Hadean eon
first ocean
Archean eans
origin of life
Proterozoic eon
first photosynthetic organisms; first eukaryotes
Cambrian
explosion of diversity of multicellular forms
ordovician
radiation of marine invertebrates, first land plants
silurian
first vascular plants
devonian
first seed bearing plants; first jawed fish; first arthropods
carboniferous
large swamps of coal
permian
giant insects and amphibians; diverse ray finned fish
triassic
conifers dominant land plant; frog and reptile diversity
jurassic
dinosaurs; pterosaurs and rayed finned fish diversity, first flowering plant
cretaceous
flowering plant diversity, animals diversity on land sea
tertiary
radiation of mammals; climates cool and grasslands increase
quaternary
evolution of homo
mesozoic era includes
triassic, jurassic, cretaceous
Cenozoic era includes
tertiary, quaternary
Paleozoic era includes
cambrian, ordovician, silurian, devonian, carboniferous, permian
what eons are in the hanerozoic
Paleozoic, mesozoic, cenozoic
alveolates
unicellular, key feature being alveoli, phototrophic lineages being dinoflagellates, secondary endosymbiosis, heterotrophic lineages are ciliates, and apicomplexans
alveolates lineages
dinoflagellates, apicomplexans, ciliates
stramenopiles
uni and multicellular, key feature: 2 unequal flagella, phototrophic is diatoms and brown algae, heterotrophic oomycetes, secondary endosymbiosis
stramenopiles lineage
diatoms, oomycetes
rhizarians
unicellular, key feature: produce shells, phototrophic some cercozoa, heterotrophic foraminifera, radiolarians, rest of cercozoa, secondary endosymbiosis
rhizarian lineages
cercozoa, foraminifera, radiolarians
excavates
unicellular, have flagella, phototrophic: euglenoids, heterotrophic: parabasilids, diplomonads, secondary endosymbiosis
excavate lineages
euglenids, parabasilids, diplomonads
plants
uni and mulitcellular, key feature: photosynthetic, phototrophic lineages: red and green algae, land plants, primary endosymbiosis
amoebozoans
unicellular, key feature: amoeba like with pseudopods, heterotrophic: loboseans, plasmodial and cellular slime molds
groups of plants
bryophytes, pteridophytes, gymnosperms, angiosperms
bryophytes key groups
liverworts, mosses, hornworts
all bryophytes have
stomata (except liverworts), need water for fertilization, gymnosperm
pteridophytes key groups
club mosses, horsetails, ferns
all pteridophytes have
stomata, need water for fertilization, and roots, sporophyte
gymnosperm key groups
conifers, ginkgoes, cycads
all gymnosperms have
stomata, tracheids and roots, pollen, seeds, sporophyte
angiosperm key groups
flowering plants
all angiosperms have
stomata, tracheids and roots, pollen, seeds, double fertilization, sporophyte
key eukaryotic traits
flexible cell membrane (lost cell wall), complex cytoskeleton, nuclear membrane, digestive vacuoles, endosymbionts (mitochondria and chloroplast), sexual reproduction (mitosis and meosis)
what are the 8 major groups of eukarya?
alveolates, stramenophiles, rhizarians, excavates, plantae, amoebozoans, fungi, animals
which lineage of eukarya does not have nuclei?
excavates lineages: diplomonads and parabasalids
2 hypotheses for the origin of mitochondira
late origin, excavates split off before the origin of mitochondria, early origin, mitochondria arose at beginning but lost in excavates
What is proven through evidence?
single origin of mitochondria before excavates separate because genes found in nuclear genomes
excavates are usually
anaerobes and are parasites
how did secondary endosymbiosis occur:
green algae: transferred to the ancestor of euglenids and chlorarachniophytes
red algae: transferred to the ancestor of the chromalveolates
ancestrally how did organisms replicate
binary fission
steps for binary fission
replication of DNA from parent cell —> segregation of DNA —> cell splitting into 2
in binary fission what is shared
plasmids which contains genes for antibiotic resistance or pathogenicity
haploid
only one of each chromosome
diploid
two of each chromosome
sex
exchange of genetic material
reproduction
making more cells
meiosis
division of 1 diploid cell to generate 4 haploid cells
fertilization
fusion of 2 haploid gametes to form the diploid zygote
diplontic life cycles
organism is diploid and the gametes are the only haploid stage
haplontic life cycle
mature organism is haploid and the zygote is only diploid stage
haplo-diplontic life cycle
organism spends time in both haploid and diploid stages
isogamy
the gametes are the same size
anisogamy
two gametes are different sizes
oogamy
egg is larger and sperm is smaller
is both gametes are mobile it is best to be
isogamous
some asexual reproducing organisms can turn sexual reproducing during times of stress because
mating locus encodes a transcription factor driving gene expression
which of the following is a key eukaryotic trait that facilitated cell movement?
flexible cell membrane, complex cytoskeleton
How do we know that excavates once had mitochondria but lost them?
they have homologs to proteobacterial genes in their nuclear DNA, they have hydrogenosomes with double membranes
in chlamydomonas how does the mating type gene impact gamete formation?
expression of the gsp allele makes a positive mating type
paramecium have an unusual sexual cycle. Which of the following is the most similar to what you would expect for orgnaism’s that have more normal sexual reproduction?
the 2 cells exchange one haploid micronucleus
What is the main difference between the 2 different kinds of slime molds in their vegetative state?
plasmodial slime molds have many nuclei per cell, the cellular slime molds have one nuclei per cell
how does the mating type locus affect the zygote
expression of the gsp and gsm alleles makes a zygote which develops, and undergoes meiosis
in what eon was their visible life and fossil evidence for multicellular organisms
phanerozoic
during which period do 75% of all animals go extinct and the sea levels drop by 50m
silurian
multicellularity
cooperation among cells for “good” of organisms
five requirements for multicellularity
extracellular environments, division of labor, resource allocation, proliferation inhibition, programmed cell death
multicellularity evolved multiple times because
it is profitable for organisms
type of multicellular designs related to nutrition
photosynthetic, adsorptive organisms, ingestive organisms (animals)
convergent designs for photosynthetic organisms
flat leaves, tubular stems with roots
ingestive organisms have specialized tissues:
muscular- to move, nervous- to coordinate, digestive- to ingest
4 major multicellular clades
land plants, animals, fungi, brown alage
closest unicellular relative to land plants
green algae: charophytes
closets unicellular relative of animals?
choanoflagellates
closet unicellular relative for fungi
opisthokonts
closet unicellular relative for brown algae
unicellular stramenopiles (diatoms)
why would a choanoflagellate benefit from being reversible colonial
only spontaneously form rosette colonies in presence of bacteria
fungal bodies
mycelia, networks of branche hyphae
benefits of multicellularity
overcoming long distance diffusion, task division
multicellular organisms can reproduce by
budding (asexual) and producing a single celled spore or gamete (sexual)
gsp allele
encodes transcription factor for making + proteins in the + gamete
gsm allele
encodes transcription factor for making - proteins in the - gamete
low population size induces the expression of
gsp allele
the mating type locus encodes the integrates genetic machinery that
transformed cells into 2 gamete types, enable syngamy, and drives development of zygote, enables meiosis to form 4 haploid spores
do organisms need to have sex organs to have sex
no
what an organism needs to have to have sex:
form haploid gametes, union of haploid meiotic products to produce diploid, meiosis to form haploid stage again
most animals have what kind of sexual cycles?
diplontic sexual cycles
most unicellular organisms have what kind of sexual life cycle?
haplontic life cycle
how do multicellular organisms switch between sexual and asexual reproduction?
during warm months reproduction is parthenogenetic = cloning, during cold months or limited food, sexual morphs are made which produced resistant “resting eggs”
what is the surest form of evidence that life existed during a particular geological era?
find mineralized body parts that have fossilized
which types of organisms have a division of labor with regards passing down the genetic material
a diploid organisms with some cells undergoing meiosis
closets relatives to metazoans
choanoflagellates
likely sequence of events for the evolution of multicellularity starting from unicellularity
colonial cells —> linear filamentous cells —> branches filamentous cells —> parenchymatous cells
Where were animals fossilized following the Cambrian radiation 540 years ago
burgess shale
Why was it easy for plants to live in aquatic environment
bathed in nutrients, extensive internal transport not necessary, gametes offspring transmitted by water, no desiccation, supported against gravity
colonization of terrestrial environments:
early: plants evolved a new monophyletic lineage with novel adaptations for surviving on land. later: plants made terrestrial environment more survivable existing lineage of animals able to colonize