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plants are in nested clades
a set of organisms that share a common ancestor
plantae =
red algae and green plants
have plastid derived from primary endosymbiosis
green plants=
green algae and land plants
have modified plastid, a chloroplast
land plants
have adaptations for living on land - cuticle, pores or stomata
cuticle - waterproofing
must have gas exchange
pores in upper surface of thallus probably are the earliest structure for gas exchange
true stomata originate later, perhaps more than once
all exhibit alternation of generations
Embryophytes
land plants, multicellular embryo, protect embryo w sterile tissue and retained on mother plant
alternation of generations
sporic meiosis
alternate btwn haploid and diploid generations
meiosis = spores
mitosis = gametes
happens for land plants and some green algae
zygote
first cell of the sporophyte (2n) phases
sporophyte
diploid generation that produces spores
spore
haploid reproductive cell that can grow into an adult without the fusion of another cell - wall of sporopollenin
produced by meiosis
gametophyte
haploid generation that produces gametes
gamete
haploid reproductive cell
fuses together to form a zygote
produced by mitosis
syngamy
fusion of gametes to form a zygote
Bryophytes
non-vascular land plants
hornworts, mosses, liverworts
sporophytes is attached to, and is dependent on the gametophyte
embryo is retained by the gametophyte
vascular plants
have specialized conducting tissue
vascular tissue - xylem and phloem
support - increased height above ground
distribution of water and nutrients
Seed-free Vascular Plants
ferns and lycophytes
reproduce via spores - not seeds or pollen
have independent gametophytes and sporophytes
Seeded Plants
gymnosperms and angiosperms
have pollen and seeds
gametophytes are short lived and the female is retained on the sporophytes in the seed
gymnosperms
relatively small number of species but very diverse
ginkgo, cycads, gnetales, conifers
ginkgo
up to 80ft tall, deciduous w yellow foliage
temperate, resistant to pollution so common street tree
fan shaped leaves
cynads
first fossil evidence for 280mya
most common - cycas, zamia, macrozamia
restricted to tropical and warm temperate regions
gnetales
ephedra - 40 species
gnetum - 30 species
welwitschia - 1
deser plant with two leaves that continue to grow and split
conifers
sporophytes - tall,wide trees or shrubs, none herbaceous
monopodial growth, most evergreen, few deciduous
male gametophyte = pollen grain
female gametophyte = inside the ovule - becomes the seed
are the largest, the oldest, and the tallest living organisms
angiosperms - monocots -
cotyledons - 1
vascular tissue - scattered through stem
veins - parallel veins in leaves, bundles of vasc tissue
flowers -petals in multiples of 3
angiosperms - eudicots -
cotyledons - 2
vascular tissue -circular arrangement through stem
veins - branching veins in leaves
flowers -petals in multiples of 4 or 5
membranes - lipid bilayers
from when lipid molecules are aligned in paired sheets
hydrophilic heads - interact w water
hydrophobic tails - interact w one another
selective permeability
small, uncharged molecules cross membranes easily
ions, large molecules diffuse slow or not at all
assisted by membrane proteins
phospholipids
move within the membranes
in constant lateral motion, rarely flip to the other side of the bilayer
permeability’s relation to fluidity
higher temperatures increase fluidity
diffusion
spontaneous movement of molecules and ions
solutes move randomly in all direction
solute
dissolved substances
passive
no input of energyc
concentration gradient
difference in solute concentrations across the space
net movement away from regions of high concentration of the solute
occurs within fluids, gasses, and across membranes
equilibrium
molecules randomly distributed throughout the solutes
but the movement of the solute never stops
can occur during diffusion
Rate of Diffusion
F = kA ((∆c) / d)
F = rate of diffusion
k = diffusion constant (depends on solute, membrane, temperature)
A = surface area for diffusion
∆c = change in concentration
d = distance for diffusion (e.g., membrane thickness)d
diffusion w charged particles
charged particles rarely cross a phospholipid membrane on their own
must take into account charge and concentration
concentration and electrical gradient
electrochemical gradient
diffusion of charged particles depending on combo of concentration and electrical gradient
electrochemical equilibrium
combo of concentration and electrical gradient are balanced
consequence of size and shape - getting bigger
increased distance between surface and center
decreases surface area to volume ratio
increased diffusion distance - substances reach center at a lower rate
decreased surface area to volume ratio means
if surface area is smaller in relation to the volume then there is less surface through which diffusion can occur
decreasing SA:V
solution - become long and thin or flat to offset effects of increased size
vacuole
reduce internal volume
in plant cells they reduce the volume of the protoplast
osmosis
special diffusion case w water
semi-permeable membrane
only unbound water can cross the membrane
free water moves from regions of low solute concentration to high concentration - this changes volume and solute concentration
low concentration of solute = high conc of free water
high conc of solute = low conc of free water
semi-permeable membrane
allows water to cross the membrane but doesn’t let most or all solutes cross
polar
molecule is bent, carries a partial negative charge on one end and partial positive on the other
hypertonic
water flows out of the vesicle - shrinks
hypotonic
net flow of water into the vesicle, swell or burst
isotonic
no change
passive transport
when substances - ions or molecule - move across the plasma membrane in absence of an outside energy source
occurs along electrochemical gradient - doesn’t require energy expenditure
channel proteins are selective
carrier proteins
undergo a change in shape to carry a molecule
conformational change
selective
each protein only permits a particular type of ion or molecule to pass through
forming a pore to allow movement through the membrane
aquaporin
allow water to move across membrane but exclude other molecules and ions
gated channels
open in response to a signal
ex - voltage gated K+ channel
active transport
cells can move molecules in a directed manner or against the electrochemical gradient
requires energy investment
uses pumps
secondary active transport
gradient is used to move a molecule or ion of interest
these pumps move material against their concentration gradients and establish different electrochemical gradients
gradients power movement of another molecule against its gradient - atp not directly used to power, electrochemical gradients power
cotransport
pump
membrane protein that provides active transport of molecules across the membrane
proton pump H+ ATPase
uses atp to move protons H+
often important in change to pH and creating electrochemical gradient
sodium-potassium pump - Na+/K+ - ATPase/K+-ATPase)
uses atp
transport Na+ and K+ against their concentration gradient
cotransporters - symporter
transport solutes against their concentration gradient using energy released by the transport of another molecule moving in the same direction along its concentration gradient
cotransporter - antiporter
transport solutes against their concentration gradient using energy released by the transport of another molecule moving in the opposite direction along its concentration gradient
tree of life 3 domains or 2?
bacteria 2.archaea 3.eukaryotes
or
bacteria 2.archaea and eukaryotes
eukaryotes
cells tend to be larger than bact + arch with more organelles and more extensive cytoskeleton
have nuclear envelop - membrane barrier separating nucleus from cytoplasm in eukaryotic cells
multicellularity has evolved many times - rare in b + a
reproduce asexually via mitosis or sexually via meiosis
protists
nott monophyletic but paraphyletic
all eukaryotes except land plants, fungi, and animals
eukaryotes lineage split bya and lots of divergence
no synapomorphies
protists just 10% of species but super abundant
paraphyletic
a group containing a common ancestor and some of the descendants
crop diseases
irish potato famine caused by protist Phytophthora infestans - water mold
plankton
diatoms and other small organisms that drift in the open ocean or lakes
form bottom of food chain - nutritional relationship among organisms
primary producers
species that produce chemical energy by photosynthesis
photosynthesis
conversion of light energy from the sun into chemical energy - carbohydrates - used by organisms
energy is stored in tissues as starch, sugar, and cellulose - wood, coal, gas, oil
stored energy can be released by burning or metabolism
source of oxygen in the atmosphere
photosynthesis equation
6CO2 + 6H2O = C6H12O6 + 6O2
light energy is used to reduce carbon, removes hydrogen from water and adds it to carbon
carbon
essential element of all life forms
carbon sinks
long - lived carbon repository from marine protists
sedimentary rocks
from protist shells made of calcium carbonatep
petroleum
accumulation of dead bacteria, archaea, and protists at the bottom of the ocean
respiration
takes sugars and metabolizes them for energy
carried out by all living organisms
mitochondria
all eukaryotes have them or genes normally found in mito
organelle that generates ATP using pyruvate as the electron donor and oxygen as the ultimate electron receptior
semi - autonomous
semi-autonomous
organelle replicates via fission, have own ribosomes and manufacture some of own proteins, have own genome - arranged as circular molecular as in bacteria, double membrane, similar in size to proteobacteria
ATP stands for
adenosine triphosphate
theory of endosymbiosis
mitochondria originated when a bacterial cell was taken up by a host cell
Lynn Marguis finally accepted in 1970
most profound evidence - orgin of the mitochondria is the close relationship between the mitochondrial genome and bacterial genomes - used lateral gene transfer
endo
inside
sym
together
bio
living
plastid
also hypothesized to have originated by endosymbiosis
protist - multicellularity
arose independently in many different lineages
allows specialization
not all cells can express the same genes so it allows for an organisms body to built in many ways
slime molds
cells can live independently part of the time but also make up a cohesive multicellular reproductive structure
protists structures provide
support and protection
foraminiferans - make up the white cliffs of dover
protists obtaining food
can be autotrophic or heterotrophic
ingestive feeding, phagocytosis, absorptive feeding, photosynthesis
autotrophic protists
can synthesize their own complex organic compounds from inorganic sources
heterotrophic
must digest organic compounds for nutrients - including parasites
ingestive feeding
eating live or dead organisms or organic debris
phagocytosis
engulf food
novel method for securing nutrients take in food much larger than single molecules
absorptive feeding
nutrients are taken up directly from the environment
usually cross the plasma membrane using transport proteins
can be parasitic or decomposers
protist - mobility
many move actively to find food
amoeboid motion, flagella, cilia
amoeboid motion
pseudopodia streams forward and rest of the cell follows
flagella
a long cellular projection that undulates - in eukaryotes to push a cell through and aqueous environment
dinoflagellates
have 2 flagella
create red tides - huge population growth = blooms
fish, bird, turtle, manatee, and dolphin death
neurotoxic shellfish poisoning - harm people who eat shellfish
also toxin in aerosois - breath it in
opisthokonta
have single flagellum
cilia
short, filamentous projections that move the cell or circulate fluids
life cycle
sequence of events that occur as an individual grows, matures, and reproduces
asexual
any mechanism of producing offspring that doesn’t involve the production of and fusion of gametes
based on mitosis - in eukaryotes
chromosomes in the daughter cell are identical to chromosomes in the parental cell
sexual
the production of offspring through the production and fusion of gametes
requires meiosis - reduced the number of chromosome sets
offspring have chromosome complements different than both parents
life cycles affects
relative timing of mitosis and meiosis affects the overall lifecycle
mitosis facilitates growth of a multicellular body
meiosis and syngamy change chromosome numbers
syngamy
union of gametes
fertilization is a type of syngamy - union of a flagellate sperm and a non-motile egg
haploid
1 set of chromosomes