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growth
an irreversible increase in volume or dry weight (biomass) of living cells
can be measured as increase in fresh or dry weight or in volume, length, height, or surface area
plant development
progress through the plant life cycle
seed germination, growth of vegetative organs and tissues
initiation and maturation of reproductive organs and tissue
fertilization, seed development and maturation
senescence and death
photosynthetic active radiation
(PAR)
400 to 700 nm
heliotropic
movements are used by some plants
leaf angles are adjusted throughout the day
maintain 90 degree angle if incidence to increase light
photomorphogenesis
highly integrated plant growth and developmental processes in response to light
de-etiolation
greening of young plants
stem growth
typically, presence or absence of light
may have more specific light spectral composition requirements and changes in leaf structure.
most photomorphogenic responses regulated by
phytochrome
phytochrome
pigment that has 2 interconvertible forms
red light absorbing
far-red light absorbing
cryptochromes
pigments involved in photomorphogenic responses and circadian rhythm
phototropin
a blue light receptor is responsible for phototropism
photoperiodism
the photomorphogenic response to variations in day length
all photoperiodically controlled processes can be categorized into 3 basics types
photoperiod plants
long day
short day
day neutral
critical day length (CDL)
flowering (other processes) induced only at day lengths longer/shorter than specific day length
stratification
some seeds require time during which the seeds are imbibed at low temps before germination is possible
chilling requirement
the duration required to break dormancy
vernalization
cold induction of flowering
determinate growth
after vegetative growth, flower bud clusters form at shoot terminals so mot shoot elongation stops
indeterminate
plants bear flower clusters laterally along the stem in the axils of the leaves
annuals
complete life cycle in one growing season
biennials
complete life cycle requires 2 growing seasons
perennials
remain alive indefinitely
annuals life cycle
germination
vegetative growth
reproductive growth
flower
fruit
seed
death (senescence)
biennials life cycle
limited stem growth 1st season
dormant
reproductive growth 2nd season
perennials life cycle
herbaceous types: shoots may die in winter, new shoots emerge in spring
woody types: root and shoot systems remain alive
shoots may go dormant in winter
juvenility
vegetative only
maturation
reproduction possible
phases of development
juvenility
maturation
senescence
death
circadian rhythms
biological rhythms that complete one cycle in approximately 24 hours
anthesis
flower opening
self pollination
same flower
same plant, different flower
different flowers, same clone
cross pollination
different plant/clone
self fertile
plant produces fruit and seed with its own pollen
self sterile
plant requires pollen from another plant
incompatibility
cross pollination between 2 cultivars can be ineffective
factors inhibiting pollen tube germination or elongation
parthenocarpy
fruit formed without pollination and fertilization
stenospermocarpy
fertilization but embryo aborts
no viable seed is produced
certain seedless grapes
simple sigmoid
slow start
period of rapid size increase
decreased rate near fruit maturity
double sigmoid
single sigmoid growth curve is repeated
climacteric fruits
as they ripen experience a burst of respiration and release high levels of ethylene
ex. apples, apricots, bananas
senescence
terminal, irreversible deteriorative change in living organisms, leading to cellular and tissue breakdown and death
plant hormone
natural chemicals that acts to control plant activities (produced by itself)
plant growth regulators
compounds applied to affect growth/development
plant growth hormone classes
gibberellin
auxin
cytokinin
ethylene
abscisic acid
gibberellin (GAs)
stimulate cell division, cell elongation or both and can control enzyme secretion
auxin
seed germination
apical dominance
one of the most widespread auxins that occurs naturally in plants INDOLEACETIC ACID (IAA)
cytokinin
primarily promote cell division
ethylene
induces fruit ripening
abscisic acid
regulation of processes in seed development (seed dormancy)
a mobile stress hormone in which ABA action initiates responses to cold and water stress
signal guard cells to close stomate and conserve water
secondary hormones
salicylic acid
jasmonates
systemin
salicylic acid
response to pathogen attack
assonates
defense against herbivores
systemin
signaling peptide that induces defense genes
basis for plant nutritional needs
carbs
lipids and fatty acids
proteins
starch
main plant stored carb
cellulose
main structural component in plants
essentially insoluble in water
glucose
6 carbons
most important and common of the sugars
fructose
5 carbon sugar
fruit sugar
sucrose
glucose + fructose
table sugar
unsaturated fatty acids
2 or more double or triple bonds between carbons (places for more hydrogens)
in human diet considered to be more healthy
saturated fatty acids
only single bonds present between carbons
in human diet considered to be less healthy
triglycerides
three fatty acids linked to glycerol
waxes, fats, oils
phospholipids
glycerol molecule with a phosphate group and 2 fatty acids
secondary products
alkaloids
phenolics
terpenoids
alkaloids
bitter taste and acts as detterents
alkaloid examples
morphine
cocaine
nicotine
caffeine
phenolics
flavanoids
anthocyanins
flavonoids
act as pigments in flowers to attract insects and birds
absorb UV light
anthocyanins
antioxidant activity
helps plant cells deal with external stresses such as cold snaps and excess light
thought to be a health component of our diet
terpenoids
rubber (latex)
carbon cycle
starts with photosynthesis (capture of solar energy)
ends with respiration (release of energy)
carbon
forms strong, stable bonds
transfer of electrons (energy) between molecules
oxidized compounds (respiration)-easily accept electrons
reduced compounds (PS)-easily donate electrons to other compounds
photosynthesis equation
6CO2+6H2O=C6H12O6+O2
PS
process of converting oxidized CO2 to reduced C
RS
process of converting reduced carbon to oxidized carbon to generate other forms of energy that are central to the function of living organisms
types of photosynthetic pigments
chlorophyll a and b
carotenoids
xanthophylls
chlorophyll a
most important pigment
absorbs blue, red, and violet wavelengths
carotenoid
yellow to orange
xanthophyll
yellow
light reactions location
grana and thylakoid of chloroplast
light rxn product
ATP and NADPH
2 phases of light rxns
photolysis of H2O
phosphorylation
dark rxns location
stroma
calvin cycle steps
carbon fixation
reduction phase
carb formation
regeneration of RuBP
classification of plants
C3 unregulated
C4 concentration regulated
CAM time regulated
C3 pathway C fixation
CO2 attached to 5C RuBP molecule resulting in 6C molecule
6C splits into 2 3C molecules (3PGA)
reaction accelerated by RuBP carboxylase (rubisco)
CO2 fixed as part of carbohydrate
C reduction
utilizing ATP and NADPH 3PGA is reduced to G3P
C3 pathway carbohydrate formation
G3P makes fatty acids, glycerols, glucose phosphate, fructose, starch, cellulose, amino acids
C3 RuBP regeneration
RuBP used in CO2 fixation must be replaced
every 3 turns of calvin cycle 5 G3P (3C MOLECULE) USED
to remake 3 RuBP (5C molecule) require ATP
C4 pathway
high light intensity, high temp, low moisture
mesophyll cells
PEP- a carboxylase PS in low [CO2]
fix CO2 to PEP (3C)
oxaloacetate (4C)
calvin cycle occurs where in C4
bundle of sheath cells
crassulacean acid metabolism
CAM
during night CAM
stomata open
fix CO2 to form 4C molecules
during day CAM
stomata closed (H2O conservation)
NADPH and ATP available
4C molecules release CO2 to calvin cycle
respiration
extracts energy stored in glucose
converts it to other forms (ATP) to perform metabolic tasks
requires carbs, oxygen and ADP
produces 36 ATP
step 1 respiration
glycolysis
conversion of glucose to pyruvate
takes place in cell cytoplasm
process of glycolysis results in
2 pyruvic acid+2ATP+2NADH
RS 2nd step is called
tricarboxylic acid (TCA) cycle
takes place in cell mitochondria
several rxns involve acids with 3 carboxyl groups