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hearing, balance, and gestation(taste)
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sensory neural adaptation
A short-term decrease in sensitivity to a constant stimulus, a type of auditory adaptation
evolutionary adaptation
A long-term genetic change in a species to improve survival and reproduction, a type of auditory adaptation
pinnae
the external part of the year, also called external auditory meatus
pinnae are directional
the trait of the pinnae that aids in localization, and makes it highly mobile
spectral filter
pattern of ridges inside pinnae act as a ______ , emphasizing some frequencies and de-emphasizing others.
middle ear
made up of the tympanum and the ossicles
tympanum
membrane that seals ear canal, vibrating when struck by sound waves from ear canal, converts sound energy into kinetic energy(referred to as eardrum)
ossicles
a series of tiny bones forming articulated chain, connects the vibrating of the tympanum to the inner ear.
malleus, incus and stapes
the 3 tiny bones that make up the ossicles, can be described as the hammer, anvil and stirrup
eustachian tube
this is used to maintain equal air pressure in ear, connects the middle ear to the oral-nasal cavity which allows bacteria to get in
tensor tympani
a small muscle that connects the malleus(ossicle) to the tympanum, a middle ear muscle
stapedius
a muscle that connects the stapes(part of ossicle) to the floor of the middle ear
tensor tympani and stapedius
these two muscles connect at each end of the ossicles to contract and give volume control to respond to sounds,
frequency
on a sine sound wave, the time from peak to peak, in Hz/CPS
amplitude
peak height(loudness) measured in dyn/cm2 or db
harmonics
multiples of the fundamental frequency of an emitter
timbre
the unique “signature” sound of an emitter, comprised of the fundamental frequency, overtones and harmonics.
doppler shift
occurs if the emitter is in motion, velocity is added to the rarefaction-compression cycle to change frequency, used by bats
resonance
there is a certain frequency for each space that is the loudest of all frequencies for that specific place. the volume of space that you’re in determines which frequency will be reinforced.
inner ear
the organ that actually neurally encodes the sound elements, known as a transducer
the cochlea
the lobsided croissant that is filled with endolymph and perilymphs, and connects to the ending of the ossicles at the oval window.
action of the stapes
vibrations transmitted from tympanum are communicated to the endolymph via the _____________ against the oval window
round window
the _____ bulges to accommodate pressure when the wavelike movements of the endolymph move through the length of the cochlea
the organ of corti
the most important part of the cochlea for hearing, consisting of the basilar membrane, tectorial membrane and hair cells.
basilar membrane
on the base of the cochlea;one of the membranes that divides the tubes of the cochlea
tectorial membrane
hangs above the organ of corti on the cochlea; another divider of the cells of the tubes of the cochlea
inner hair cells
when stimulated, they release glutamate(excitatory) on to auditory nerve fibers. the base of them is near the basilar membrane, and the tips grace the tectorial membrane.
stereocilia
sensory “antennae” of the ear. They bend with sound, triggering electrical signals. Crucial for turning mechanical sound energy into nerve impulses
3500
how many IHC
12000
how many Outer Hair Cells(OHC)
Outer hair cells
same as stereocilia, but releases Acetylcholine, and is influence by GABA.
amplitude of the travelling wave
for any given frequency, the ___ __ ___ ________ is exaggerated on one point of the basilar membrane
Afferent IHC
leads to brain. carries the info that we perceive as sound. activated by glutamate from the IHC
Efferent IHC
from brain to IHC. Serve a modulatory function by inhibiting the IHC-afferent fibres. Uses Acetylcholine.
Afferent OHC
from OHC to brain, small diameter nerve fibres using ACh, convey activity info of basilar membrain to brain. not involved in concious perception of sound
Efferent OHC
from brain to OHC. using GABA, alters the responsiveness of OHC, tuning
tether
the tip of each stereocilium is connected to its neigbour by this tip link.
pores
when stereocilia bend, the tip links are stretched, popping open the ____ on the tethered stereocilia
instantaneous depolarization
when pores open, it is large and cations rush in to create this:
tuning
the action of the basilar membrane changing its width
neural tuning
when inputs from numerous auditory fibres converge on neuronal systems that filter our the frequency of the received sound.
lateral inhibition
when frequencies that the neurons selectively respond to selectively inhibit their neighbouring frequencies
electromechanical tuning
the voltage dependent tuning of the OHC to change the responsiveness basilar membrane
otoacoustic emissions
the active nature of the cochlea causes it to emit clicks at aroud 20db
evoked OAE
provoked by presented sounds: useful for testing hearing in infants, effects of drugs on hearing, experiments on basic cochlear mechanisms, a type of otaacoustic emissions
spontaneous OAE
unnoticeable otoacoustic emissionsm associated with sensitive hearing, females experience more, along with right ear(left hemisphere) potentially related to language
projections
the sound info that is conveyed to higher brain structures in a complex way, probably due the variety of things that produce sound info
cochlear nuclei
a major centre of projection- in the brainstem, it is the first receiving centre in the brain, it is for low level integratoin, tuning and projections to other areas(projects sounds to high order functions of brain
superior olivary complex
a major centre of projection: brainstem, receives and integrates inputs from both cochlea, basis of binaural(stereo) hearing.
inferior colliculus
a major centre of projection: midbrain, tuning, spatial localization, you can see differences in sound from left and right ear
medical geniculate
a major centre of projection: part of the thalamus, projects to auditory cortex
auditory cortex
superior temporal cortex: primary auditory area
pitch discrimination
the idea that pitch is not the same as frequency, the note A can still be heard as A even if its 1 or 2 Hz off 440 Hz
Place theory
The theory that perceived pitch corresponds to the location on the basilar membrane that is most strongly activated(you perceive pitch along the basilar membrane)
volley theory
the theory that pitch is a function of the rate of firing in auditory fibres. higher frequency = many waves per unit of time, more action potentials and vice versa
volleys
the discharge rates that are directly related to sound frequency which occur only below 1500 Hz
sound localization
this is best done by binarual earring, but can also be done with just one ear if you’re free to move your head
intensity differences
sounds off the midline result in difference in amplitude intake for each ear
head shadow
the head blocks sounds from getting to the more distant ear causing the sound intensity differences worsen
time of arrival
the angle of the sound source is directly related to the difference in arrival time at the two ears
out of phase
term to describe the peaks and troughs of the sound wave(compressions and rarefactions) have to go farther to hit the more distant ear and thus arrive a little later
direction of the pinna
this trait of your “ear meat” discriminates front from back audio
duplex theory
each separate ear’s intensity and arrival time comes together to produce a sound. this is the :
cortical deafness
cortical lesions DO not produce ______, because the auditory cortex is not all-important for hearing basic soudn
cortical areas
areas of the auditory system that are devoted to recognizing important sounds: footsteps, animal calls, vocalizations of familiar versus unfamiliar people
right auditory cortex
the side of the auditory cortex that associates to many music functions
left auditory cortex
planum temporal bigger on this side of the auditory cortex due to this side having speech specialization
conduction
the often type of deafness, a problem with the ossicles
sensorineural
most often cochlear deafness, hair cells blown over like a bunch of tiny trees, can be caused by loud soudns, certain drugs(aspiring and streptomycin)
central deafness
a type of deafness arising from brain damage
vestibular system
the mechanism for detecting changes in head position(acceleration and motion sickness)
semicircular canals
the 3 components of the vestibular system, each in a different plane. fluid filled, circulates from motion of head,
ampulla
the base of the semicircular canals, hair cells translate this movement into neural signals inside this thing
otolotihs
overlying the hair cells, and amplify the effect on the hair cells. translate to “ear stones”, made of gelatin
papillae
the bumps on your tongue
fungiform papillae
mushroom-shaped, usually one taste bud each, has multiple sensory cells per taste bud
foliate papillae
leaf/paged shape papillae on side of the tongue, has multiple taste buds per one
circumvallate papillae
big suckers at the back of the tongue, contain multiple
umami
savoury, soy, beef, gravey type taste.
olfactory
what trait, in contrary to common belief, controls your sense of taste, and diminishes it when you have a cold
salt
identify the taste: simple ion pores admit sodium ions, and are captured by a specific recepter.
sour
identify the taste: derived from acids, which release hydronium ions that block potassium channels on the tastebud cells
TRPV1
identify the taste receptor: for salty food taste, secondary salt sense
bitter
identify the taste: metabotropic-like receptor, signals poison,
T2R
identify the taste receptor: bitter taste receptor
sweet
identify the taste: a metabotropic receptor, liberating a 2nd messenger.
T1R family
identify the taste receptor family: the receptors in charge of sweet/umami taste
T1R1
both sweet and umami are received by T1R3, but which one is UMAMI only received by?
T1R3
both sweet and umami are received by T1R3, but which one is SWEET only received by?
umami
identify the taste: gluatamate receptor.
olfactory epithelium
the main sensory apparatus, forming the lining of the nasal cavity,
support, basal, and olfactory receptor cells
THE 3 major cell types in the olfactory system
olfactory dendrites
the ______ ______ are studded with chemical receptors, and appear to be metabotropic-like receptors linked to a second messenger(like gpcr)
regeneration
to adapt to chemical attacks, viruses etc. olfactory cells are constantly being replaced by converting basal cells to neurons
vomeronasal organ
the second olfactory system in humans/mammals that contains biologically important stuff like pheromones and pseudogenes(V1R’s , V2R’s , and TRP2)