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major eras
paleozoic (543-252 ma): early life
mesozoic (251-65 ma): dinosaurs and early mammals
cenozoic (65 ma-present): mammals dominate, human evolution
cenozoic periods
paleogene (65-23 ma): warm, early primates
neogene (23-2.6 ma): cooler, apes and early hominins
quaternary (2.6 ma-present): ice age, homo evolution
epochs
paleocene→first primate-like animals
eocene→first true primates
oligocene→early monkeys
miocene→apes diversify
pliocene→early hominins
pleistocene→homo spreads
holocene→modern humans
types of fossils
macrofossils: bones, teeth
microfossils: microscopic remains
trace fossils: footprints, behavior evidence
fossilization conditions
rapid burial, low oxygen, hard body parts
relative dating
stratigraphy: layers of rock accumulate over time; deeper = older
paleomagnetism: uses earth’s magnetic field reversals recorded in rocks to match a fossil layer to a known time period
biochronology: uses fossil species with known time ranges to estimate the age of a site
absolute (chronometric) dating
radiometric (e.g., potassium-argon)
luminescence (OSL; sunlight)
dendrochronology (tree rings)
convergent lines of evidence
multiple dating methods used together
key primate features
forward facing eyes (binocular vision)
reduced snout (less reliance on smell)
grasping hands/feet
postorbital bar/closure
larger brains
major hypotheses for primate evolution
arboreal: adaptations for tree living
visual predation: hunting insects → depth perception
angiosperm: fruit/flower exploitation
plesiadapiforms
primate-like but not true primates
had claws, no full eye convergence
eocene primates
first true primates
warm climate
adapids (lemur-like) and omomyids (tarsier-like)
oligocene primates
cooler climate
first anthropoids (monkeys, apes, humans)
miocene apes
traits: no tail, y-5 molars, larger brains
climate cooling → forests shrink → ape decline
important miocene apes
proconsul: early ape
sivapithecus: ancestor to orangutans
european apes (dryopithecus)
key adaptations for bipedalism
foramen magnum underneath skull, s-shaped spine, short wide pelvis, valgus knee, arched foot
why bipedalism
carrying food/tools, energy efficiency, heat regulation, foraging advantages
earliest hominins
sahelanthropus: possible earliest, small brain
orrorin: femur suggests bipedalism
ardipithecus: bipedal + arboreal
australopithecus
full bipedal but still climbed, small brains, reduced canines
a. afarensis (lucy), a. africanus
paranthropus
robust, huge teeth + chewing muscles
not ancestors of homo
homo erectus
first to leave africa, larger brain (~875cc), modern body proportions
acheulean tools, possible fire use, increased social structure
long childhood, altricial babies (dependent)
homo heidelbergensis
likely ancestors of neanderthals (europe) and homo sapiens (africa)
neanderthals
large brains (~1520cc), cold-adapted bodies, skilled hunters, used tools + fire
care for injured, possible burial practices, interbred with humans (1-2% DNA in non africans)
life history theory
energy → growth, maintenance, reproduction
humans = slow life history
big brain tradeoffs
brain = 2% body weight, 20% energy
tradeoff with gut (expensive tissue hypothesis)
brain evolution hypotheses
social intelligence, behavioral flexibility, cognitive buffer