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Outer Segment (Photoreceptors)
contains photo-pigment; catches the light
Inner Segment(Photoreceptors)
Contains mitochondria; helps power the eye
Number of cones per eye
6-8 million
Number of rods per eye
120-130 million
Geographic distribution of cones
Mostly in fovea
Geographic distribution of rods
Mostly in periphery
Acuity of cones
Great
Acuity of Rods
bad
Acuity
smallest spatial detail that can be resolved at 100% contrast
Color in cones
They can see it
Color in rods
they can not see it (achromatic)
Peak wavelength in cones
About 550 nm
Peak wavelength in rods
About 500 nm
Purkinje Shift
Photopic (Day)= reds are brighter than greens; Mesopic= Greens are brighter than reds; Scotopic (Night)= greens turn into dark grey and reds turn into black
Motion in cone
Very sensitive
Motion in rods
not so sensitive
Luminance in cones
Day; not so sensitive; "photopic"
Luminance in rods
Night; High sensitivity to light; "scotopic"
Dark adaption in cones
quick (about 5 minutes)
Dark adaption in rods
slow (about 30 minutes)
Dark adaptation
Below your average threshold of what you're used to, you will not be able to see
Your pupil constricts to reduce the amount of light arriving at your retina
Time Course
the speed and the time spent traveling in a known direction from a known position, rather than a direct observation
Rhodopsin
Found in the rods of the retina; Extremely sensitive to light and therefore allows for good vision in low-light conditions
"Kinked" function on adaption graph
When the cones and rods are the most sensitive at any given point
Two parts of pigment
The protein and vitamin A derivative
Protein (ospin)
Found in the photoreceptor cells of the retina; Mediating the conversion of a photon of light into an electrochemical signal
Vitamin A derivative
actually catches the light
Transduction
changing physical energy into neural energy
Photo-isomerization
chromophore changes shape and releases energy when photon is absorbed; changes in flow of electrical current surrounding the photoreceptor occurs; takes <1 msec
Sign inverting (sign-reversing) connections
when light is absorbed it hyperpolarizes; rods and cones become more negative and cells become more positive
"Backwardness of eye"
no receptors on outside of eye
order of cells in the retina
photoreceptors--> horizontal --> bipolar--> amacrine--> retinal ganglion cells
Retinal Ganglion cells processing (RGC)
Light--> retinal ganglion cells--> amacrine cells--> bipolar cells
Two main types of RGC
Parvo-cellular (P-cells) and Magno-cellular (M-cells)
lateral inhibition
when neurons fire, they inhibit neighboring cells
Mach bands
differences in edges; darker on the dark side and lighter on the light side; exaggerating the edges
Microelectrodes and single cell recording
Microelectrodes are used in measuring single cells; they look for ways they get excited (EX: MRIs)
Optic Chiasm
x-shaped structure formed at the point below the brain where the two optic nerves cross-over each other
Temporal retina
visual track on the out side of the eyes and stays on the same side (Ipsilateral)
Nasal retina
visual track on the inside of the eyes; cross over; left nasal retina processes in the right hemisphere of brain and visa versa (Contralateral)
Superior Colliculus characteristics
Older; used to find where, not what
Layered multi-sensory structure
Superior Colliculus functions
the upper layer receives visual signals from the retina; lower layer processes multiple signals from various other parts of the brain
Jay and Sparks Superior Colliculus study
Found auditory cells in the Superior Colliculus; found that they fire when you aren't looking at where the noise is coming from and do not fire when you are look at where the noise is coming from ("Annoying little brother cells"
Lateral Geniculate Nucleus (LGN) characteristics
Small, ovoid, ventral projection at the terminal of the optic tract on each side of the brain; all about organization
Layers of LGN
6 layers- Layers 1 & 2 (bottom layers) are Magnocellular; Layers 3, 4, 5, & 6 (top layers) are Parvocellular
Magnocellular layers of LGN
Large; rods; necessary for the perception of movement, depth, and small differences in brightness ; Rapid and transient
Parvocellular layers of LGN
Small; Cones; long and medium wavelength; necessary for the perception of color and form (fine detail); Slow and sustained
LGN functions
Receives information directing from ascending retinal ganglion cells via optic track and from the reticular activating system; Receives many strong feedback connections from the primary visual cortex; Main connection for the optic nerve to the occipital lobe
Topographical mapping
The orderly mapping of the world in the lateral geniculate nucleus and the visual cortex
Primary Visual cortex (V1) or the Striate Cortex
The area of the cerebral cortex of the brain that receives direct inputs from the lateral geniculate nucleus, as well as feedback from other brain areas
Orientation tuning
the tendency of neurons in striate cortex to respond optimally to certain orientations and less to others
Filter
an acoustic, electrical, electronic, or optic device, instrument, computer program, or neuron that allows the passage of some range of parameters and blocks the passage of others
Ocular Dominance
The property of the receptive fields of striate cortex neurons by which they demonstrate a preference, responding somewhat more rapidly when a stimulus is presented in one eye than when it is presented in the other
Simple Cells
A cortical neuron whose receptive field has clearly defined excitatory and inhibitory regions
Complex cells
A cortical neuron whose receptive field does not have clearly defined excitatory and inhibitory regions
End Stopping
The process by which a cell in the cortex first increases its firing rate as the bar length increases to fillip its receptive field, and then decreases its firing rate as the bar is lengthened further
Column
A vertical arrangement of neurons. Neurons within a single column tend to have similar receptive fields and similar orientation preferences
Hyper-column
A 1-millimeter block of striate cortex containing two sets of columns, each covering every possible orientation, with one set preferring input from the left eye and one set preferring input from the right eye
Cytochrome Oxidase (CO)
An enzyme used to reveal the regular array of "CO blobs," which are spaced about 0.5 millimeter apart in the primary cortex
Cortical Magnification
The amount of cortical area (usually specified in millimeters) devoted to a specific region in the visual field
Perceptual consequences of Cortical Magnification
Visual Crowding- The deleterious effect of clutter on peripheral object recognition
Schneider's Experiment
Used hamster to investigate if the midbrain was involved with eye movement. They used hamsters because their brains aren't fully developed and they are easily manipulated with sunflower seeds
What did the Schneider experiment find
Focal vision was used to identify what and ambient vision was used to determine where
Differences in focal and ambient vision
retinal location; awareness (ambient needs it); Luminance (focal requires a lot); refractive error (Ambient is okay with some); optical rearrangement; spatial frequency
Civil Twilight
The time between day and night when the sun is below the horizon but its rays still light up the sky (brightest of the three phases)
Civil Twilight times
About 30 minutes before sunrise and about 30 minutes after sunset
Applied vision with driving
Uses both focal and ambient visual processes
Andre & Owens ciil twilight distance
the distance in front of the vehicle where the dark limit of civil twilight is reached and focal vision can be considered useful
How far down the road is the dark limit of civil twilight
225 feet at ground level (can vary)
Spatial Frequency Analysis
The number of cycles of a grating per unit of visual angle (usually cycles per degree)
Cycles per degree
The number of pairs of dark and bright bars per degre of visual angle
On-Center receptive fields
A cell that depolarizes in response to an increase in light intensity in its recepetive-field center; Wants the light in the center of the receptive field and dark on the outside (chocolate donut)
Off-Center receptive fields
A cell that depolarizes in response to a decrease in light intensity in its receptive-field center; Wants it dark in the center and light on the outside
Contrast
The difference in luminance between an object and the background, or between lighter and darker parts of the same object; represented by the waves amplitude
Fourier Analysis
Mathematical procedure by which any signal can be separated into component sine waves at different frequencies. Combining these sine waves will reproduce the original signals (Square waves are made up of a ton of separate sine waves)
Sine-Wave Graph physiology
Different cells = different size receptive fields = different spatial scare
Receptive fields
the region on the retina in which visual stimuli influence a neuron's firing rate
What are Sine-wave gratings
Visual stimulus with alternating darker and lighter areas with a sinusoidal luminance profile (gradual transitions into light and dark)
Sine-Wave characteristics
Higher Specific frequencies = thinner strips (finer detail)
Color vision cones
S-cones; m-cones; l-cones
S-Cones
peak at 420 nm
M-cones
peak at about 535 nm
L-cones
peak at about 565 nm
Principle of Univariance
the fact that an infinite set of different wave length intensity combinations can elicit exactly the same response from a single type of photoreceptor
Hue
divides blues, reds, greens, etc
Brightness
Related to the amount or intensity of light (lightness or darkness of the color); the more light absorbed the darker the surface will appear
Saturation
Amount of hue; pale or vivid
Color spindle
Circumference = hue; radius = saturation; height = bightness
Young-Helmholtz trichromatic receptor theory
a theory that the color of any light is defined in our visual system by the relationships of three cones (S,M,L); a "lower" theory = the actual sensation
Opponent Processing theory (Hering)
color is from the output of three mechanisms, each of them resulting from an opponency between two colors (red-green, blue-yellow, and black-white); a "higher" theory = where the processing occurs
Complementary colors
Orange and blue, Yellow and Purple, red and green
Additive color mixing
Light; approaching white when you mix
Subtractive color mixing
Pigments; The typical idea of mixing blue and yellow to get green
Color Vision Deficiencies in gender
More males than females have color deficiencies
Achromatopia
Missing all cones
Abnormal Trichromats
Red-Green Protanomaly, Red- Green Deuteranomaly
Dichromats
red-green protanipia, red-green deuteranopia, blue-yellow tritanopia
Red-Green Protanomaly
Abnormal L pigments
Red- Green Deuteranomaly
Abnormal M pigments
Red-green protanipia
Missing L pigments