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life without touch
trouble standing, walking, getting burned; very rare
somatosensory system
touch: pressure, temperature, pain
proprioception: perception of own body; sense of movement and position of limbs
the skin
receptor organ for touch and pain
epidermis
outer layer of the skin
dermis
layer beneath epidermis
mechanoreceptors
receptor cells responding to mechanical stimulation (pressure, stretching, vibration)
4 types of mechanorecpetors
Merkel receptor, Meissner corpuscle, Ruffini cylinder, Pacinian corpuscle
adaption rate
fast adapting and slowly adapting
fast adapting (FA)
fire only at the onset and offset of stimulation; Meissner and Pacinian
proprioception
perception of your own body; sense of movement and position of the limbs
properties of mechanoreceptors
adaptation rate, detail resolution
detail resolution
size of receptive field
merkel receptor
sensitive to fine detail
pacinian receptor
sensitive only to larger grain
primary somatosensory cortex (S1)
receives touch information from the opposite side of the body; each point varies in size
somatotopic map
organization of the primary somatosensory cortex maintaining a representation of the arrangement of the body.
tactile acuity
ability to detect details on the skin
two-point threshold
minimum separation between two points on the skin that when stimulated is perceived as two points
grating acuity
placing a grooved stimulus on the skin and asking the participant to indicate the orientation of the grating
temporary cause of loss of touch sensation
limbs "falling asleep", numbness from cold
permanent cause of loss of touch sensation
damage to brain or spinal cord due to accident or stroke, diseases
phantom limb
when an amputee has the sense that an amputated limb is still attached to their body
two components of pain
responses of pain receptors, cognitive and affective aspects
nociceptors
several types of receptors in skin that are sensitive to extreme heat, cold, pressure, and chemical exposure
gate control theory
the spinal cord contains a neurological "gate" that blocks pain signals or allows them to pass on to the brain
3 input pathways to T cells
S-fibers, L-fibers, central control fibers
S-fibers
carry pain signals from nociceptors
L-fibers
carry info about non-painful tactile simulation
central control fibers
carry signals about cognitive function like expectation, attention, and distraction
Rainville et al. (1997)
manipulated perceived unpleasantness of stimuli (hot water)
Eisenberger et al. (2003)
social pain; participants reporting greater distress → greater activation of ACC
analgesia
inability to feel pain
hyperalgesia
increased sensitivity to pain
haptic perception
the active exploration of the environment by touching and grasping objects with our hands
exploratory procedure
stereotypical hand movement pattern used to contact objects in order to perceive their properties
taste
sensations produced when various substances dissolved in saliva penetrate the taste buds on the tongue and surfaces of the mouth
gustation
technical name for taste
flavor
combination of sensations, including food's taste, smell, temperature, texture, and consistency
5 basic tastes
sweet, sour, salty, bitter, umami
papillae
bumps on tongue
taste buds
cluster of elongated cells in throat, roof of mouth, inside of cheeks (although not inside papillae)
taste receptors
respond to dissolved compounds
microvilli
small threads at end of receptor cell
tongue map
debunked theory of where specifically certain receptors for taste lie on the tongue
specific taste receptors
particular taste receptor cells for particular tastes
tasters
people who can taste PTC
nontasters
people who cannot taste PTC
supertasters
individuals with heightened taste sensitivity
primary taste cortex
identifies and judges intensity of tastes
orbitofrontal cortex
combine with other senses and enables pleasure from food
Rolls (2004)
at first, much neural firing for smell of cream but after eating 50ml, smell produces less firing
Mozel et al. (1969)
smell improved identifying food greatly when tasting it
retronasal olfaction
the perception of odor caused by chewing food in the mouth
specific hungers theory
the idea that deficiency of a given nutrient produces craving for that nutrient
labeled lines theory
each taste fiber carries a particular taste quality
Penfield & Rasmissen (1950)
discovered S1 as somatotopic map; stimulate point on S1 and ask what’s perceived
Singer et al. (2004)
empathetic pain; shock → S1 + ACC, seeing shock → ACC
Melzack & Wall (1965)
gate control system
touch pathway
skin → nerve fibers → spinal cord → ventral posterior nucleus → cortex
anterior cingulate cortex (ACC)
region of the brain associated with perceived unpleasantness of pain sensation
tactile agnosia
inability to identify objects by touch
slow adapting (SA)
fire continuously as long as pressure is applied; Merkel + Ruffini
high acuity
more Merkel receptors + corresponding space in S1