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evolutionary biology
historical science because it is figuring out what has happened over millions of years
two ways of reconstructing evolutionary history
phylogenetics and the fossil record (can get lots of info with these two together)
phylogenetics
living species and how they are related to each other to figure out evolution
fossil record
looking at past life
phylogenetic tree
diagram that describes phylogeny
phylogeny
geneology on broadest possible scale of living things
why is the phylogenetic a hypothesis
still constantly collecting data to find the evolutionary relationships among various species
parts of phylogenetic tree
branches and nodes
branches
represent populations
nodes
hypothetical ancestors that gave rise to two species
two axes of phylogenetic tree
time and some kind of change
what does a phylogenetic tree show
speciation events and normal events of isolation and divergence
what do nodes represent
a speciation event; the common ancestor from which the species diverged
sister groups
share a common ancestor not shared by any other group; on either side of a node
monophyletic group
common ancestor and all of its descendants
paraphyletic group
common ancestor and some, but not all, of its descendants
polyphyletic group
does not include a common ancestor; just have similar characteristics
taxonomy
hierarchal system involving successive levels
taxon
group at each level of hierarchal system of taxonomy
domain
most inclusive group of taxonomy, all of life belongs to one of three domains
domains
bacteria, archae, eukarya
classification
phylogenies reflect the hierarchial taxonomic system of Linneaus; all species grouped into single phylum
homology
similarity due to common descent
analogy
superficial similarity, often due to convergent evolution
building a phylogenetic tree
comparing homologous characters by scoring character states
convergent evolution
evolution on two different species that look similar to survive in their environment
character states
what does a character look like in different groups of origins
apomorphy
derived character state
synapomorphy
shared derived character state; taxa that have synapomorphies are related by common descent
apomorphy vs synapomorphy
apomorphy is an evolutionary novelty and a synapomorphy is an apomorphy shared by two or more groups
out group
taxon that derived prior to lineage of taxa of interest
character matrix
compare homologous characteristics to put in tree
determining the best fit of possible phylogenetic trees
the best tree is the most parsimonious one/one with the fewest changes in it because it has had the least amount of evolutionary events
occams razor
simplest explanation
benefits of using molecular data to build a phylogenetic tree
vast amounts of data
fossils
only record of extinct forms
using fossils to create phylogenies
allow us to calibrate phylogenies in terms of time
provide a record of extinct species
place evolutionary events in context with Earth’s history
evolutionary events and fossils
appearance of features can be correlated with geological features
what rocks do fossils typically form in
sedimentary rocks
what makes the fossil record biased
1) common things
2) hard parts
3) aquatic
4) more recent
5) more widespread
do we have a fossil record of everything that has ever lived
no because it is not easy to fossilize
trace/molecular fossils
not body parts
evidence of animal activity (footprints, burrows, etc)
molecular seen if look closely at chemical of rocks because there are traces of biochem that is only through living things
burgess shale
some of the earliest animal fossils preserved in Canadian Rockies (early evidence of life)
soft parts preserved
messel shale
german fossils
rare
impressions
hadean era
earth on fire
archean era
first evidence of life
proterozoic era
oxygen present in earth
phanerozoic era
visible life appears
radioactive decay
computes how many half lives have passed in rock
pangea
super continent that existed 250 million years ago
what caused pangea to break up and what happened when it did
extreme environmental conditions and it led to mass extinction
mass extinction
5 large mass extinctions
boundaries between geological time periods are often based on the events
rapid extinction of species