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Chapter 4, 6, 8, 7, 11, 12
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Species Diversity
species richness + species evenness
Species Richness
The number of different species in community
Example:
Forest with 20 different species = high richness
Species Evenness
How evenly individuals are distributed among species
Example:
4 species with 25 individuals each = high evenness
1 species with 90 individuals, others with 2–3 = low evenness
Biome
large geographic area recognized
by their distinct structures Ex. biotic & abiotic factors, rainfall & and temps
*can be terrestrial or aquatic
Latitude
Def. distance from the equator
* Ecosystems near the equator have much more diversity!
○ More physical space in the tropics
○ More sunlight
○ Relatively stable climate
○ We don’t totally know why!
Equilibrium Model of Island Biogeography
The number of species
on an island is the result of a balance
between the rate at which new species
arrive and species on the island become
locally extinct
* Shaped by:
○ Island size = Larger target - more immigrants & less extinction because can maintain larger pop.
○ Distance to shore = more immigrants from close proximity - leads to less extinction because sustained by rate of immigration
Keystone Species
Exert significant
effect on biodiversity despite low
abundance. Usually a top-level predator Ex. Wolve
Ecosystem Engineer
Manipulate the
physical environment and influence
ecosystem structure. Ex. Beaver
Foundation Species
Create the physical
framework for the community using
their own bodies Ex. Trees = provide shelter and home for birds
Succession
Gradual change in a
community over time Ex. Wild flowers growing on side of mountain after volcanic erruption
Primary Succession
succession on a
bare geological surface, such as a lava
flow
○ Slow, takes decades to get fresh soil
Pioneer Community
First community to develop during succession. Generally made of species tolerant to exposure of full sun and harsh conditions
Secondary Succession
succession following the disturbance of an established community that doesn’t
destroy all living creatures or soil
Climax Community
The community at
the end of a successional sequence that
persists until a disturbance disrupts it
sufficiently to restart succession
Ecosystem Services
1. Supporting: Foundational services like
energy flows and nutrient cycles
2. Provisioning: Products obtained from
the ecosystem
3. Regulatory: Processes derived from
ecosystems
4. Cultural: Non-material benefits
○ Recreation, tourism, aesthetic inspiration
Isolated Patches
Definition:
Areas of habitat that are separated from other similar habitats by human development or natural barriers, limiting movement of organisms between them.
Why It Matters:
Reduces gene flow
Lowers biodiversity
Increases risk of local extinction
Ex.
Small forests separated by highways or cities
Habitat Fragmentation
Definition:
The process where a large, continuous habitat is broken into smaller, isolated patches.
Causes:
Roads
Cities
Farming
Dams
Effects:
Creates isolated patches
Reduces gene flow
Lowers biodiversity
Increases edge effects
Edge Effects
Changes that happen at the border of a habitat, often harming species that need the interior environment.
Caused By:
Habitat fragmentation
Creation of isolated patches
Examples of Edge Effects:
More sunlight & wind
Higher temperatures
More predators & invasive species
Lower survival of interior species
Why It Matters:
Harms specialist species
Reduces biodiversity
Makes ecosystems less stable

Protected Areas
Geographically defined area that is
designated or regulated as damaged to
achieve a particular conservation
objective Ex. Sichuan Giant Panda Sanctuary
*Strict nature reserve= No tourism specifically for scientific research only
*Size (bigger is better, but not always possible) & connection/ continuous zone (linkage of land substantiable habitat to increase movement of wildlife) is key

Non-Gov & Ngo role on conservation
Not all protected areas are run by the
government!
* NGOs have more flexibility compared to the government
Ex. World Wildlife Fund
Gene Def.
Stretch of DNA that directs the
growth, development, and functioning of
an organism
Genetic Diversity
Different gene
combinations found within a single
population or species Ex.
Natural Selection
Those with more
favorable genetic traits survive better
than others
Domestication
The deliberate change
of a wild animal or plant species to
better meet the needs of humans Ex. Dogs & bananas
*problem = threat to biodiversity Ex. bananas only having one type now because before there were more types but are vulnerable to disease - now may have to create a new variation to sustain our demand
J-Shaped (exponential) Growth
As long as growth
remains fixed, the population size will
increase faster and faster over time until crash (die and run out of resources)

S-Shaped (logistic) Growth
Limited essential resources can cause the rate of growth to slow down and eventually level off at
carrying capacity
Carrying Capacity (K)
the number of individuals in a population that an environment can support over the long run
Life History
Characteristics of a species
and the rate at which young survive
K-Selected Species
○ Regulated by density dependent factors
○ Few offspring & parental intensive care
Ex. Whales & Elephants - pop. is effected and controlled by limited food and space, competition, and use of natural resources
R-Selected Species
○ Rapid growth when the environment is
favorable
○ Small and subject to catastrophic
mortality from density independent
factors
○ Lots of offspring, little parental
investment
○ Live fast, die young!
Ex. Insect - pop. is effected and controlled by pesticides, temp changes, and natural disasters
Threats to Biodiversity #1 Habitat Destruction
We live in the Anthropocene def. a time period where humans shape the planet more than natural processes
● Habitat alteration and destruction are
the most serious threats to biodiversity
Ex. Loss of sea ice habitat for polar bears, seals, and penguins
Threats to Biodiversity #2 Invasive Species
When introduced to a new environment, pose a serious threat
to native populations
● Not hindered by their own natural
predators, parasites, and pathogens
● Happens when humans bring species to new places
Ex. Zebra mussels in the Great Lakes – outcompete native mussels and clog water infrastructure.
Threats to Biodiversity #3 Plant and Wildlife Trafficking
Species killed or removed from the
environment for human use
○ Demand for exotic pets
○ Flashy hunting trophies
Ex. Rhinos, elephants, exotic birds
Threats to Biodiversity #4 Pest and Predator control
Predators seen as threats to livestock
are killed without considering effect on
ecosystems
● Pest control introduces toxins that
harm species
Ex. Weeds- herbicides used in agriculture
Solutions to biodiversity threats- The Endangered Species Act
1- The Endangered Species Act protects
both domestic and foreign endangered
species
Solutions to biodiversity threats- The Lacey Act
2- The Lacey Act forbids the trading of
illegally harvested plants and animals
Solutions to biodiversity threats (Suggestions)
- Prevent spread of invasive species
- Ban toxins responsible for endangering species
Solutions to biodiversity threats- Economic Incentives
Encourage ecotourism
Regulated hunting
Drugs produced from different plants
Distribution of Water on Earth
1. Oceans = 97% of all water
2. Freshwater mostly in polar ice caps
3. Next largest = groundwater
4. Very little freshwater in lakes, rivers, wetlands
Frozen Water
1. Glaciers
Losing mass since 1970s
95% near the poles
Melting → sea level rise & ocean current disruption
2. Sea Ice
Mostly seasonal (~15% of oceans)
Melting does not add water
High albedo → reflects sunlight, cools Earth
Less sea ice → warmer oceans → affects global climate
Hydrologic Cycle
def. The movement of Earth’s water between the oceans, atmosphere,
and terrestrial and freshwater environment

Connection between oceanic and terrestrial sub cycles (Hydrologic Cycle)
1. Ocean → land: precipitation falls on land
2. Land → ocean: surface runoff flows back to the ocean
Water Use
We use >50% of accessible runoff water globally
Main uses:
Irrigation
Diluting sewage/waste
Rivers for shipping
Recreation
Lakes & Ponds (lentic)
Standing water
Classified by distance from shore & sunlight penetration
Rivers & Streams (lotic)
Flowing water
Classified by flow differences
Wetlands & estuaries
Wetlands = water-saturated land
Estuaries = freshwater + saltwater meet
Dams- Pros vs. Cons
Pros:
Emission-free power (hydropower)
Regulate water supply (store water during droughts)
Cons:
Rivers dry up downstream
Reduce nutrient flow below dam
Change water temperature
Block fish migration → affects aquatic biodiversity
Dam removal- Pros vs. Cons
Term: Dam Removal
Pros:
Restore natural ecosystems
Reestablish fish migration and river flow
Cons:
Lose clean hydropower
Ecosystem recovery not guaranteed
Release trapped pollutants downstream
Floods & Wetlands
Floods = rivers/streams overflowing banks
Wetlands need floods to thrive
Critical in dry water-scarce areas for biodiversity
Human changes to flooding harm wetlands
Erosion- Def. + problems
Def. =The removal and movement of soil or rock by wind, water, ice, or gravity.
Problems= Humans speed up erosion by:
Removing water / lowering water tables
Deforestation → no roots to hold soil
Groundwater Depletion
Pumped faster than recharge
Some areas use fossil water (non-renewable aquifers)
Harms groundwater-dependent ecosystems
Low flow, warmer streams → hurts salmon & trout
Ex. Ogallala Aquifer
Subsidence
def. The sinking of land when groundwater-filled spaces collapse.
Caused by overpumping groundwater
Permanent loss of aquifer storage capacity
Leads to land sinking, infrastructure damage, and reduced water supply
Irrigation
Waterlogging
Soil becomes too saturated → roots can’t get oxygen
Soil Salinization
Water evaporates and leaves salt behind
Salt buildup harms crops and reduces soil fertility
Why water rights are hard to define
Hard to measure needs - daily water needs vary by environment, age, weight etc.
Debate: water = human right vs. commodity
UN recognizes the right to water (2010), but many countries abstained
Legal/political problems:
Who pays to move water across borders?
Should countries share water with growing neighbors?
How much water is each person guaranteed?
Water reclamation
Definition:
Treating wastewater so it can be reused or recycled.
Treatment Levels & Uses:
Primary:
No reuse
Secondary:
Irrigate orchards/vineyards
Wetland restoration
Industrial uses
Tertiary:
Irrigate food crops
Landscaping
Recreational water (ponds, lakes)
Add to surface reservoirs for drinking supply
Regulations keeping freshwater clean
1. Safe Drinking Water Act (SDWA)
Sets maximum contaminant levels for drinking water
Protects water at the tap
2. Clean Water Act (CWA)
Regulates pollution discharge into water bodies
Protects rivers, lakes, wetlands from contamination
Thermohaline Circulation Def. & movements
Def. Global movement of ocean water driven by temperature (thermo) and salinity (haline).
Key Points:
Warm water moves north → cools → becomes dense → sinks
Controls global climate patterns
Keeps Europe warm compared to similar latitudes (ex: Bordeaux vs. Portland, Maine)
AMOC = part of thermohaline circulation
Thermohaline collapse (if it failed)
Major Consequences:
Dramatic cooling in Europe & North America
More flooding & storms in the Pacific
Collapse of plankton (base of food web)
Climate shifts in tropics, Alaska, Antarctica
Stronger & more frequent El Niño events
Low oxygen below ocean surface → mass extinctions
El Nino
🌞Where:
Equatorial Pacific Ocean, west of South America
What It Is:
Warmer than normal sea surface temperatures
Low atmospheric pressure
Climate Effects:
Warm & dry winters in some regions
Alters global rainfall patterns
La Nina
❄ What It Is:
Colder than normal sea surface temperatures
High atmospheric pressure
Climate Effects:
Very wet winters in Southeast Asia
Warmer winters in the Southern Hemisphere
ENSO (El Nino southern oscillation)
Def. The natural climate cycle between El Niño & La Niña
*It is a normal swinging pattern, not always caused by climate change
Upwelling
Def. When cold, nutrient-rich water rises to the ocean surface
Importance:
Supports plankton & fisheries
Cold surface spots can signal circulation changes
Marine Primary Production
Def. Energy production by phytoplankton through photosynthesis
Controlled By:
Ocean temperature
Ocean salinity
Ocean nutrients
Euphotic vs Disphotic
Euphotic Zone: Enough light for photosynthesis
Disphotic Zone: Some light, no photosynthesis
Upwelling + Fisheries
Why It Matters:
Feeds phytoplankton
Supports productive fisheries
El Nino + Fisheries
What Happens:
Warm surface water moves in
Disrupts winds & upwelling
Nutrients can’t reach euphotic zone
Results:
Commercial fish populations drop
Seabird & marine mammal deaths
Why human whaled + Consequences
Uses:
Meat
Blubber → oil, makeup, soap, margarine, industrial oil
Why It Became a Crisis:
Bigger boats → easier killing
Mostly in international waters → no laws
Led to population collapse
Regulation- International Whaling Commission (IWC)
Created to set hunting quotas
1982: Commercial whaling banned
Many whale populations are now recovering
Why whaling still continues
Cultural traditions
Indonesia, Alaska, Canada
Commercial / “scientific” whaling
Japan left IWC (2019)
Iceland still permits whaling
Tragedy of the commons- Fisheries
Why Overfishing Happens:
Non-excludable = anyone can fish
Rivalrous = less fish for others
Individuals act for personal gain
Solution 1- Private Property
Old rule: short season → race to fish
New rule: guaranteed share of catch or area
Encourages long-term sustainability
Solution 2- Pigouvian Tax
Fishing licenses limit who can fish (but easily given out)
Economic control to reduce overuse
Small-scale but somewhat effective
Marine Reserves- Types + Ex.
Definition:
Ocean areas with restricted or banned fishing
Types:
Permanent no-take
Seasonal closure
Benefit:
Fish populations recover & spill over
Ex. Cabo Pulmo
Overfished in 1990s
Became no-take reserve
Fish biomass increased ~5× in 10 years
Coral reef recovery
Boosted eco-tourism & local economy
Aquaculture
Definition:
Breeding & raising fish, shellfish, algae for food
Why It’s Used:
Rising food demand
Rapid industry growth
Land-based aquaculture
def. Fish raised in indoor tanks with filters
Ex.
Salmon in China & Middle East deserts
Tuna in Japan
*Downside: expensive, energy & water intensive
Problems with aquaculture
Escaped fish become invasive
Water pollution from waste & chemicals
Disease & parasite transfer to wild fish
Carnivorous fish still require wild-caught feeder fish
IMTA- Integrated multi-trophic aquaculture
Def. Raising multiple species from different trophic levels together
Pros=
Waste from one species feeds another
Reduces pollution & disease
Less fish meal needed
Fewer pesticides
Aquatic-Terrestrial Linkage
Definition:
Connections between water ecosystems and land ecosystems where energy, nutrients, organisms, and carbon move between them
Key Idea:
What happens in streams, rivers, and lakes directly affects nearby land ecosystems
Riparian food web
Definition:
Food webs that exist along stream and river edges (riparian zones)
Key Connections:
Aquatic insects emerge from water → become food for land predators (birds, spiders)
Nutrients move from water → into terrestrial ecosystems
Why Important:
Streams support biodiversity on land
Human impact on riparian ecosystems
1 Pollution (pesticides, chemicals) → kills aquatic insects
2 Fertilizer runoff → changes aquatic community
3 Dams & channels → destroy insect larval habitat
4 Habitat destruction → removes riparian vegetation
Result:
Disrupts food supply for terrestrial predators
How to protect riparian ecosystems
Solutions:
Protect stream health
Increase riparian buffer zones
Restore natural water flow + vegetation
Why It Matters:
Freshwater management = protects land biodiversity too
Land + Atmosphere carbon exchange
Old View:
Land and water tracked separately
New Understanding:
Carbon moves laterally through watersheds
Carbon travels by:
Surface runoff
Groundwater flow
Carbon can be:
Released into the atmosphere
Buried in aquatic sediments
Big Idea:
Carbon cycling is land + water + atmosphere together
Measuring carbon exchange-
Top down vs Bottom up
Top-Down Method (Atmosphere Based):
Measures total carbon exchange from above
✅ Sees the big picture
❌ Doesn’t explain exact ecosystem processes
Bottom-Up Method (Land Based):
Adds together carbon from forests, lakes, farms, etc.
✅ Explains how ecosystems work
❌ Accuracy depends on good ecosystem data
Net Watershed Exchange (NWE)
Definition:
Measures total carbon balance of an entire watershed
Includes:
Carbon stored in ecosystems
Carbon exported by streams
Why Better:
Looks at whole carbon system, not just water transfer
Virtual Water
Def. The hidden water used to produce goods, especially food and crops
Example: Water used to grow cotton or cocoa
Why do water-scarce countries export virtual water?
Economic Reasons:
Comparative advantage (they make more money exporting)
National economies may depend on these exports
Example: Cotton industry
Cultural & Social Reasons:
Farming water-intensive crops is tied to:
Tradition
Social status
Community identity
Example: Volta Basin (Ghana): cocoa farming & livestock
Big Idea:
Money + culture often outweigh water shortages
Terrestrial Primary Production
3 Most Important Factors:
Climate – controls rate of photosynthesis
Biodiversity – different species fill different roles → increases stability & productivity
Nutrients (soil) – soil supplies nutrients needed for plant growth
Nitrogen importance
Nitrogen:
Essential for amino acids & proteins
Supports plant growth & food chains
Often the limiting nutrient in terrestrial ecosystems
Nitrogen Cycle
Definition:
The movement of nitrogen through and between ecosystems
Main Steps to Know:
Nitrogen fixation
Decomposition & ammonification
Nitrification
Assimilation
Denitrification
Weathering
Soil importance
Soil Is More Than Dirt
Soil Is:
A living system
Made of:
Sand
Silt
Clay
Soil Development:
Takes hundreds to thousands of years
How nutrients enters and leaves soil
Nutrients Enter Through:
Nitrogen fixation
Weathering
Deposition
Plant litter
Sedimentation
Nutrients Leave Through:
Erosion
Leaching
Gaseous emissions
Erosion
Definition:
Movement of soil from one place to another
Causes:
Farming
Construction
Deforestation
Overgrazing
Effects:
Loss of nutrients
Reduced soil fertility
Lower crop yields
Ways human get food
Foraging:
Hunting, fishing, gathering
Agriculture:
Extensive – small land, more labor
Intensive – large land, less labor, high inputs
Uses:
Plows
Draft animals
Irrigation
Fertilizers
Pesticides
Green Revolution
Time Period:
Mid-20th century
Goal:
Increase food production & reduce hunger
Key Figure:
Norman Borlaug
Results:
✅ Increased yields
✅ Reduced hunger
❌ Environmental damage
❌ Increased inequality
❌ Big corporations replaced local farms
Agroecology
Def.
Farming using ecological knowledge to increase sustainability & biodiversity
Includes:
Crop rotation
Intercropping
Low-till & no-till
Cover crops
Green manure
Organic farming
Key Idea:
Farms function like ecosystems
Agroforestry
Def.
Combining trees + crops in farming
Benefits:
Reduces erosion
Helps fight climate change
Limits deforestation
Challenge:
Must match natural species correctly
Indigenous knowledge in farming
Why It Matters:
Proven sustainable techniques
Improves:
Soil management
Crop rotation
Intercropping
Example:
Three Sisters = corn, beans, squash
Natural Farming
Definition:
Rejects modern chemical-based farming
Pros:
Very eco-friendly
Cons:
Productivity depends on high skill & experience
Example:
Ducklings in East Asia for weed & pest control
Ranching & Overgrazing
Ranching:
Raising livestock
Problems:
Requires large land & energy
Causes overgrazing
Overgrazing Effects:
Up to 100 tons of soil lost per hectare per year
Leads to desertification
Example:
Sahel region
Forest management practice #1
Clear-cutting
Most economical
Removes all trees
High erosion