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weather
short term variation in temperature and precipitation (hours or days)
climate
long term typical atmospheric conditions through the year (measured over many years)
example of events that can be rare but have large effects
tsunamis
example of some variation occurring in regular intervals
forest fires
large-scale spatial variation
impacted by factors like climate, land topography, and soil type
small-scale spatial variation
impacted by factors like plant structure and animal behavior
the faster an individual moves through space and the smaller the scale of spatial variation the ?
shorter the temporal scale of variation
all phenotypes result from ?
genes interacting with environments
environmentally induced phenotypes have a genetic ?
basis
a phenotype well-suited to one environment may be ? suited to other environments
poorly
phenotypic trade-off
a situation in which a given phenotype experiences higher fitness in one environment, whereas other phenotypes experience higher fitness in other environments
phenotypic plasticity
the ability of a single genotype to produce multiple phenotypes
phenotypic plasticity allows organisms to achieve ? if environmental conditions vary
homeostasis
natural selection favors the evolution of ? if the environmental variation results in ?
phenotypic plasticity; phenotypic trade-offs
phenotypic plasticity example
Gray tree frog tadpoles produce a phenotype that allows fast escape when predators are present and fast growth when predators are absent.
what phenotype is favored if spatial variation is not common
single phenotype
acclimation
an environmentally induced change in an individual's physiology
slow responses to acclimation
change in morphology (body shape) and life history (time to sexual maturity)
rapid responses to acclimation are ? whereas slow responses are ?
reversible; irreversible
species alter their ? ? and ? in response to the presence of predators (adaptation)
growth, body shape, and behaviors
example of adaptation
when Virginia pepperweed is eaten by herbivores, the plant develops leaf hairs which makes the leaves difficult to consume
variation in resource example
the Burmese python may come across a meal once a month. when a rodent is consumed, the snake doubles the length of its intestine and quadruples its heart size to accommodate the extra tissue
hermaphrodites
individuals that produce both female and male gametes; individuals are able to fertilize their eggs with their own sperm (self-compatible)
inbreeding depression
the decrease in fitness caused by matings between close relatives due to offspring inheriting deleterious alleles from both the egg and the sperm
organisms cam adjust their physiology to maintain activity across environmental temperatures
example: isoenzymes in goldfish
microhabitats
locations within a habitat that differ in environmental conditions from the rest of the habitats
microhabitats example
the desert iguana regulates its body temperature by basking on rocks, seeking shade, or burrowing in the ground
if plants are scarce to water, how do they adapt to their environment?
they close their stomata or grow longer roots using energy and materials
migration
seasonal movement of animals from one region to another.
monarch butterflies example
they migrate to wintering areas in southern north america and to the north during the summer
dormancy
a condition in which organisms dramatically reduce their metabolic processes
4 types of dormancy
diapause, hibernation, torpor, aestivation
diapause
involves a partial or complete physiological shutdown in response to unfavorable conditions; common in insects
hibernation
individuals reduce the energetic costs of being active by lowering heart rate and decreasing body temperatures; common in mammals
torpor
a brief period of dormancy that occurs in birds and mammals in which individuals reduce their activity and their body temperature
aestivation
the shutting down of metabolic processes during the summer in response to hot or dry conditions
foraging
behavior associated with recognizing, searching for, capturing, and consuming food (plastic behavior)
animals have 4 responses to food variation in space and time
central place foraging, risk-sensitive foraging, optimal diet composition, diet mixing
central place foraging
foraging behavior in which acquired food is brought to a central place, such as a nest with young
central place foraging example
a nest with young birds
traveling time
the time needed to travel round-trip to site with resources
searching time
the time spent obtaining food at a site
risk-sensitive foraging
foraging behavior that is influenced by the presence of predators
sensitive foraging example
creek chub feed on tubifex worms, but locations with worms also contain more predators
optimal diet composition
most animals do not consume a single food item, and base their diet decisions on handling time in addition to the energetic and nutritional value of various resources.
handling time
the amount of time that a predator takes to consume a captured prey
diet mixing
some foragers consume a varied diet because one type of food might not provide all of the necessary nutrients.
diet mixing example
humans combine food items to consume all essential amino acids
adaptation
seemingly good fit of the organism to the environment
water is densest at
4ºC
viscosity
the thickness of a fluid that causes objects to encounter resistance as they move through it
water's viscosity is
high
solutes
dissolved substances in water
semipermeable membrane
membranes that allow only particular molecules to pass through
osmosis
movement of water across a semipermeable membrane
osmotic potential
the force with which a solution attracts water by osmosis
osmoregulation
mechanisms organisms use to maintain a proper solute balance
hyperosmotic
tissue solute concentrations are higher than surrounding water
hyposmotic
tissue solute concentrations are lower than surrounding water
sharks and rays convert ammonia into what?
urea
salt balance in plants example
mangrove trees have high concentration of organic solutes in their roots and secrete salt from their leaves
aquatic plants use bicarbonate or carbonate for what
photosynthesis
countercurrent circulation
adaptation where blood and water flow in opposite directions so that the concentration of O2 in water is always greater than the concentration in blood
anaerobic/anoxic
environment becomes completely without oxygen
thermophilic
organisms living up to 110ºC
glycerol and glycoproteins
chemicals present in some animals that prevent freezing by reducing strength of hydrogen bonds or via supercooling
thermal optima
the range of temperatures in which an organism best performs
isoenzymes
different enzymes that catalyze same reaction
coral bleaching
loss of color in corals as a result of the corals expelling their symbiotic algae
cohesion
the mutual attraction of water molecules; allows water to move up through empty remains of xylem cells
root pressure
when osmotic potential in the roots of a plant draws in water from the soil and forces it into the xylem elements
transpiration
leaves can generate water potential as water evaporates from the surfaces of leaf cells
cohesion-tension theory
the mechanism of water movement from roots to leaves due to water cohesion and water tension
stomata
small openings on leaf surfaces that are points of entry for CO2 and exit points for water vapor; bordered by guard cells that and open and close each stoma
electromagnetic radiation
energy from the sun, packaged in small particle-like units called photons
photosynthetically active region
wavelengths of light that are suitable for photosynthesis
chloroplast
cell organelles found in eukaryotic photosynthesis organisms
chlorophylls and carotenoids are pigments inside ?
the thylakoids absorb light
chlorophylls absorb ? and reflect ?
red/violet light; green/blue light
chlorophyll a is found in ? and primarily responsible for ?
all plants; photosynthesis
chlorophylls b, c, d, f are ? pigments
accessory
chlorophylls b, c, d, f capture light energy and pass it to ?
chlorophyll a
carotenoids reflect ?
orange/red light
carotenoids allow plants to absorb ?
a wider range of solar energy
photosynthesis occurs in two steps ? and ?
light reactions; Calvin cycle
Calvin Cycle takes place in the ?
stroma of the chloroplasts
the energy in ATP and NADPH is used to convert CO2 into ?
glucose
Calvin Cycle C3
most plants/ most common
uses Rubisco
inefficient
ideal for wet/cool climates
Calvin Cycle C4
grasses, sedges
derived
Pep and OAA
reactions physically separated
ideal for hot/dry climates
Calvin Cycle CAM
family Crassulaceae (stonecrops)
derived
Pep and OAA
temporal separation of reactions
ideal for hot/dry climates
homeostasis
an organism's ability to maintain steady internal conditions when outside conditions change
negative feedback
the action of internal response mechanisms that restores a system to a desired state, or set point, when the system deviates from that state
negative feedback example
in mammals, the hypothalamus triggers increased metabolism when body temperature is below 37ºC and sweating when body temperature is above 37ºC
animals acquire ? in the water and food they consume
mineral ions
water intake and urine excretion eliminate excess ?
salts
water and salt balance example
the desert kangaroo rat conserves water by hunting during the night and staying below ground during the day
organisms that do not have access to fresh water (marine iguana) eliminate the salt in their drinking water through ?
specialized secreting organs
most aquatic animals eliminate excess nitrogen as ?
ammonia radiati
mammals produce ?
urea
birds and reptiles produce ?
uric acid