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protocells
droplets wit membranes that maintain an internal chemistry different from that of their surrounding.
stromatolites
layered rocks that form when certain prokarytoes bind thin films of sediment together.
endosymbioosis
when a prokaryote cell engulfs another small cell.
serial endosymbiosis
suggests that the mitochondria evolved before plastids through a sequence of endosymbiotic events
plastids
double membrane organelles in plant and algae cells. these are responsible for many things like photosynthesis, pigment synthesis, and nutrient storage.
phylogeny
evolutionary history of a species or group of species
taxon
named group at any level of hierarchy
phylogenetic tree
evolutionary history of a group of organisms can be represented with this
branch point
represents a the common ancestor of two evolutionary lineages diverging from it.
evolutionary lineage
sequence of ancestral organisms leading to a particular descendant taxon
sister taxa
groups of organisms that share an immediate common ancestor
rooted phylogenetic tree
a branch point (often farthest to the left) represents the most recent common ancestor.
basal taxon
lineage that diverges from all other members of its group early in the history of the group
homology
phenotypic and genetic similarities due to shared ancestry
analogy
similarities between organisms due to convergent evolution
cladistics
common ancestry is used to classify organisms
clades
groups that contain ancestral species and all of its descendants
monophyletic group
consists of one ancestral species and all of its descendants
parapyletic group
contains an ancestral species and only SOME of its descendants
polyphyletic
most recent common ancestor of the members is not part of the group
shared ancestral character
characteristic that originated from an ancestor of the taxon
shared derived character
a character shared by all of the group but not found in their ancestors. evolutionary novelty unique to the clade
outgroup
species/group of species from an evolutionary lineage that is closely related but not part of the group of species being examined
ingroup
group of species being examined
orthologous genes
homology is the result of a speciation event and occurs between genes found in different species.
paralogous genes
homology results from gene duplication, thus multiple copies of these gens have diverged from one another within a species.
molecular clock
approach for measuring the absolute time of evolutionary change based on the observation that some genes and regions of genomes appear to evolve at a constant rate.
horizontal gene transfer
process in which genes are transferred from one genome to another through mechanisms such as exchange of transposable elements and plasmids, and viral infections.
prokaryotes
single celled organisms that make up the domains bacteria and archae. They can also survive in a large range of environment
peptidoglycan
what most bacterial cell walls are made of
polymer composed of modified sugars cross-linked by short polypeptides
gram stain
technique used to categorize bacteria by observing differences in cell wall composition
gram positive
relatively simple wall composed of a thick layer of peptidoglycan
gram negative
walls have less peptidoglycan and are structurally more complex
outer membrane contains lipopolysaccharides
capsule
sticky outer layer of polysaccharide protein that covers the cell wall of many prokaryotes.
is also called a slime layer if it is not well defined
this layer protects against dehydration and some can protect against attacks to the immune system
endospores
developing resistant cells during times where there is minimal water or nutrients.
fimbriae
hairlike appendage that some prokaryotes use to stick to their substrate
pili
appendages that pull two cells prior to DNA transfer from one cell to the other
taxis
directed movement towards or away from a stimulus
move towards oxygen/nutrients (positive)
move away from toxic substance (negative)
nucleoid
where prokaryotes have their chromosomes
plasmids
smaller rings of independently replicating DNA molecules that prokaryotic cells have (most carry only few genes)
transformation
where genotype and possibly phenotype of a prokaryotic cell are altered by the uptake of foreign DNA from its surroundings.
transduction
phages (bacteriophages) carry prokaryotic genes from one host cell to another.
conjugation
DNA is transferred between two prokaryotic cells (usually the same species) that are temporarily joined. always one way (bacteria)
F factor
piece of DNA responsible for for the ability to form pili and donate DNA during conjugation.
F plasmid
F factor in plasmid form
R plasmid
similar to F plasmid and often carry resistance genes (code for enzymes that specifically destroy/hinder effectiveness of certain antibiotics).
obligate aerobes
must use O2 for cellular respiration and cannot grow without it
obligate anaerobes
poisoned by O2
some will live exclusively on fermentation while others will extract chemical energy by anaerobic respiration
facultative anaerobes
use O2 if it is present but can also carry out fermentation or anaerobic respiration if there is no oxygen present
nitrogen fixation
some cyanobacteria and methanogens use this method to convert atmospheric nitrogen into ammonia. This ammonia is used in amino acids and other organic molecules
heterocysts
specialized cell in a filamentous chain that performs nitrogen fixation
biofilims
metabolic cooperation among cells of one or more prokaryotic species ocurring in surface coating colonies
extremophiles
first prokaryotes in domain Archaea that live in extreme environments.
extreme halophiles
live in highly saline environments
extreme thermophiles
live in high temperature environments
methanogens
archaea that lives in a moderate environment. releases methane as by-product of how they obtain energy
exotoxins
proteins secreted by certain bacteria and other organisms that are poisonous.
endotoxins
lipopolysaccharide components of the outer membrane of gram-negative bacteria. released only when the bacteria die and their cell walls break down.