speed trig

Speed Spanish II Lesson 2

Speed Spanish II Lesson 8

Speed Spanish II Lesson 9

Speed Spanish II Lesson 10

Speed Spanish II Lesson 11

Speed Spanish II Lesson 12

Speeding Up the Action Potential, the Neuromuscular Junction, and CNS Synapses

Plant and Seed Scientific Names

Program Design: Speed & Agility

regret doing sth – żałować, że coś się zrobiło I regret saying that – żałuję, że to powiedziałam regret not doing sth – żałować, że się czegoś nie zrobiło be in two minds about sth – wahać się; mieć mieszane uczucia co do czegoś be in sb’s best interests – leżeć w czyimś najlepszym interesie have sth in mind – mieć coś na myśli make a decision – podjąć decyzję regret a decision – żałować decyzji an argument in support of sth – argument przemawiający za czymś psychological discomfort – dyskomfort psychiczny mental torment – udręka psychiczna cause a great deal of sth – powodować bardzo dużo czegoś prevent sb from doing sth – powstrzymywać kogoś przed zrobieniem czegoś move on with your life – iść dalej ze swoim życiem get caught in a trap – wpaść w pułapkę an „if only” trap – pułapka myślenia „gdybym tylko…” at certain points in our lives – w pewnych momentach naszego życia be incapable of doing sth – nie być zdolnym do zrobienia czegoś strive to do sth – usilnie starać się coś zrobić strive as we may – choćbyśmy nie wiem jak się starali come to the right conclusion – dojść do właściwego wniosku know the outcome of sth – znać rezultat czegoś there is no way of knowing – nie ma sposobu, żeby wiedzieć based on information available at the time – na podstawie informacji dostępnych w danym momencie be kinder to ourselves – być dla siebie bardziej wyrozumiałym regardless of how things turn out – niezależnie od tego, jak potoczą się sprawy release ourselves from regret – uwolnić się od żalu an empowering thing to do – coś, co dodaje siły i sprawczości regardless of + noun / -ing – bez względu na + rzeczownik / czasownik z -ing notwithstanding + noun – pomimo + rzeczownik (formalnie) otherwise – w przeciwnym razie; inaczej nonetheless – niemniej jednak; mimo to have sth in common – mieć coś wspólnego What do X and Y have in common? – Co X i Y mają wspólnego? even if – nawet jeśli (warunek, możliwość) even though – chociaż; pomimo że (fakt) not have the faintest idea – nie mieć bladego pojęcia the clue lies in sth – wskazówka / rozwiązanie tkwi w czymś a pairing of names – zestawienie dwóch nazwisk a double act – duet występujący razem be known as sth – być znanym jako coś comedy duo – duet komediowy perform as one – występować jako jedność either … or … – albo … albo … on stage / on screen – na scenie / na ekranie do so – zrobić to; postąpić w ten sposób for their entire working lives – przez całe życie zawodowe form a close bond with sb – nawiązać bliską więź z kimś establish themselves as – ugruntować swoją pozycję jako a mainstay of TV entertainment – filar rozrywki telewizyjnej set up a joke – przygotować grunt pod żart less defined roles – mniej wyraźnie określone role odd-sounding – dziwnie brzmiący such a success – taki sukces remind sb of sth – przypominać komuś o czymś whatever the reason – niezależnie od powodu be here to stay – zadomowić się na dobre one of the performers – jeden z występujących other / another / the other – inni / kolejny, inny / ten drugi both … and … – zarówno …, jak i … secret languages – tajne języki undeniably – niezaprzeczalnie be around for a long time – istnieć od dawna deliberately – celowo; świadomie play on sb’s ignorance – wykorzystywać czyjąś niewiedzę communicate with a confidante – porozumiewać się z powierniczką utterance / utterances – wypowiedź / wypowiedzi develop early in childhood – rozwijać się we wczesnym dzieciństwie begin to develop – zacząć się rozwijać what later becomes a language – coś, co później staje się językiem it may seem illogical – może się wydawać nielogiczne communicate in a language different from … – porozumiewać się w języku innym niż … mother tongue – język ojczysty an insignificant number – nieznaczna liczba as a natural consequence of sth – jako naturalna konsekwencja czegoś at the same rate – w tym samym tempie mislead sb – wprowadzić kogoś w błąd understand each other – rozumieć się nawzajem without intervention – bez interwencji language development – rozwój językowy acquire a language – przyswoić język occasionally – od czasu do czasu into adulthood – aż do dorosłości toddlers – małe dzieci uczące się chodzić language input – bodźce / informacje językowe, które dziecko otrzymuje inability to do sth – niezdolność do zrobienia czegoś psychologically and linguistically – psychologicznie i językowo no deliberate attempt to do sth – brak celowej próby zrobienia czegoś caregivers – opiekunowie reinforce errors – utrwalać błędy cause setbacks – powodować opóźnienia / niepowodzenia have no difficulty in doing sth – nie mieć trudności z robieniem czegoś continue to use – nadal używać despair that … – rozpaczać, że …; tracić nadzieję, że … have no sense of direction – nie mieć orientacji w terenie whatsoever – w ogóle; jakikolwiek (dla podkreślenia) you are not alone – nie jesteś sam(a) a significant number of people – znaczna liczba osób find sth problematic – uważać coś za problematyczne navigate / navigating – nawigować / poruszać się; orientować się w terenie be off the beaten track – być poza uczęszczanymi szlakami be concerned about sth – martwić się czymś use sat-navs – korzystać z nawigacji satelitarnej refer to maps – korzystać z map / odwoływać się do map ask for directions – pytać o drogę find sth intimidating – uważać coś za onieśmielające avoid going to unfamiliar locations – unikać chodzenia w nieznane miejsca an intriguing new type of belt – intrygujący nowy rodzaj paska Bluetooth connection – połączenie Bluetooth when worn – kiedy jest noszony indicate which direction is north – wskazywać, gdzie jest północ be armed with a device – być wyposażonym w urządzenie relieve the demands on the eyes and ears – odciążyć oczy i uszy guide sb through vibrations – prowadzić kogoś za pomocą wibracji locate / pinpoint a place – zlokalizować / dokładnie wskazać miejsce take sb’s focus off sth – odwrócić czyjąś uwagę od czegoś around their waists – wokół ich talii track / path / way / road – szlak / ścieżka / droga / droga-ścieżka the potential of an innovation – potencjał innowacji make sth more straightforward – uczynić coś prostszym a decade or so ago – około dziesięciu lat temu sporting events – wydarzenia sportowe originally – pierwotnie sports teams – drużyny sportowe catchy chants – chwytliwe okrzyki acquire different associations – nabrać innych skojarzeń a sport in its own right – sport sam w sobie; pełnoprawna dyscyplina clubs are springing up everywhere – kluby pojawiają się wszędzie at an all-time high – na rekordowo wysokim poziomie a unique combination – wyjątkowe połączenie acrobatics – akrobatyka an additional sport – dodatkowa dyscyplina sportowa in addition to teamwork – oprócz pracy zespołowej participants point out that … – uczestnicy wskazują, że … improved fitness – lepsza kondycja fizyczna safety measures – środki bezpieczeństwa be thought to be risky – być uważanym za ryzykowne join a squad – dołączyć do zespołu be associated with sth – być kojarzonym z czymś come about – powstać; wydarzyć się a form of encouragement – forma zachęty synchronised dance routines – zsynchronizowane układy taneczne be echoed by the crowds – być powtarzanym przez tłumy popularity as a sport – popularność jako dyscyplina sportowa rapidly increasing – szybko rosnący competition entries – zgłoszenia do zawodów have its own distinctive style – mieć własny charakterystyczny styl group gymnastics – gimnastyka zespołowa a touch of glamour – odrobina blasku demand strength and coordination – wymagać siły i koordynacji solid teamwork – solidna współpraca zespołowa the benefits of getting involved – korzyści z zaangażowania się a sense of camaraderie – poczucie koleżeństwa; wspólnoty be considered paramount – być uważanym za najważniejsze bear this in mind – mieć to na uwadze hot desking – praca przy dowolnym dostępnym biurku flexible working hours – elastyczne godziny pracy spend quality time with family – spędzać wartościowy czas z rodziną impose a policy – narzucić politykę / zasady maximise the availability of sth – maksymalizować dostępność czegoś the practice has been in place for years – praktyka obowiązuje od lat questionable popularity – wątpliwa popularność contain personal belongings – zawierać rzeczy osobiste opt to accept sth – zdecydować się coś przyjąć beat the rush-hour traffic – uniknąć korków w godzinach szczytu it would not be unreasonable to … – nie byłoby nierozsądne, żeby … compelling reasons – przekonujące, ważkie powody office equipment – sprzęt biurowy establishments – instytucje / przedsiębiorstwa dedicated office space – własna, przypisana przestrzeń biurowa be deeply rooted in sb – być głęboko zakorzenionym w kimś primeval fear – pierwotny lęk fear of being unsettled – strach przed brakiem stabilizacji wanderer / traveller – wędrowiec / podróżnik have no base – nie mieć stałej bazy feel undervalued – czuć się niedocenianym bring a definite element of freedom – wnosić wyraźny element wolności not be given personal space – nie otrzymywać własnej przestrzeni lack of dedicated space – brak przydzielonego miejsca have / has / had – mieć; czasowniki pomocnicze czasów perfect not only + auxiliary + subject + verb, but … – nie tylko …, lecz także …; inwersja na początku zdania Not only did he love rock bands, but he was also a fan of classical music. – Nie tylko kochał zespoły rockowe, lecz był też fanem muzyki klasycznej. concede that + clause – przyznać, że … The director conceded that he had underestimated the power of the situation. – Dyrektor przyznał, że nie docenił powagi sytuacji. as though / as if + clause – jak gdyby; jakby He looked as though he hadn’t slept all night. – Wyglądał, jakby nie spał całą noc. be thought to have + past participle – uważa się, że ktoś/coś zrobił(o) coś w przeszłości The vegetable is thought to have been brought here. – Uważa się, że warzywo zostało tu sprowadzone. point out that + clause – wskazać / zwrócić uwagę, że … My manager pointed out that I had made many mistakes. – Kierownik wskazał, że popełniłam wiele błędów. had + past participle + before + past simple – czynność wcześniejsza przed inną czynnością przeszłą I had completed three questions before I realised my mistake. – Ukończyłam trzy pytania, zanim zdałam sobie sprawę z błędu. mistake – błąd We must get to the bottom of what is causing the leak. – Musimy ustalić, co powoduje wyciek. get to the bottom of sth – dotrzeć do sedna / przyczyny czegoś We had a discussion about how to approach the problem. – Dyskutowaliśmy o tym, jak podejść do problemu. have a discussion about sth – przeprowadzić dyskusję na temat czegoś Our parents were appreciative of our efforts. – Rodzice doceniali nasze wysiłki. be appreciative of sth – doceniać coś; być wdzięcznym za coś Our parents took notice of our efforts. – Rodzice zauważyli nasze wysiłki. take notice of sth – zwrócić uwagę na coś Our products will still be enjoyed in years to come. – Nasze produkty nadal będą cieszyć klientów w przyszłości. will still be + past participle – nadal będzie + imiesłów bierny; konstrukcja strony biernej We heard about the rescheduling of the event. – Usłyszeliśmy o zmianie terminu wydarzenia. hear about sth – usłyszeć o czymś My exam results exceeded my expectations. – Wyniki egzaminów przekroczyły moje oczekiwania. exceed sb’s expectations – przekroczyć czyjeś oczekiwania even if – nawet jeśli (warunek) even though – chociaż (fakt) although / though – chociaż; mimo że while – podczas gdy; chociaż; w czasie gdy as – jako; ponieważ; kiedy; tak jak (zależnie od zdania) as though – jak gdyby such – taki (np. such a success – taki sukces) whom – kogo; komu (formalnie) whether – czy (alternatywa / niepewność) whatever – cokolwiek; bez względu na to, co however – jednak; jakkolwiek therefore – dlatego; zatem otherwise – w przeciwnym razie regardless of – bez względu na in spite of / despite – pomimo (+ rzeczownik lub -ing) in addition to – oprócz; dodatkowo (+ rzeczownik lub -ing) about – o; około into – do środka; w (np. get into) out – na zewnątrz; część czasowników frazowych (np. point out) off – z dala; od (np. off the beaten track) still – nadal; wciąż already / yet – już / jeszcze; już (w pytaniach i przeczeniach) be / been / being – być / III forma czasownika „be” / forma -ing be in two minds ABOUT (not: of) – wahać się co do czegoś prevent sb FROM + -ing – powstrzymywać kogoś przed zrobieniem czegoś regret + -ing (past action) – żałować czegoś z przeszłości opt TO + verb – zdecydować się coś zrobić be concerned ABOUT – martwić się o coś be known AS – być znanym jako be thought TO BE – być uważanym za have difficulty IN + -ing – mieć trudności z robieniem czegoś allow sb TO DO sth – pozwolić komuś coś zrobić point AT sth – wskazać palcem na coś hear OF sb/sth – słyszeć o istnieniu kogoś/czegoś look AS THOUGH + clause – wyglądać, jak gdyby … in its OWN right – samo w sobie; niezależnie come ACROSS (find by chance) – natknąć się na coś set UP (organise) – zorganizować; założyć get TO the bottom OF – dotrzeć do sedna bear IN mind – mieć na uwadze incapable OF + -ing – niezdolny do zrobienia czegoś avoid + -ing (not: avoid to do) – unikać robienia czegoś regret to inform (formal news) – z przykrością informować be associated WITH – być kojarzonym z be linked TO – być powiązanym z be a fan OF – być fanem be thought TO HAVE DONE – uważa się, że ktoś coś zrobił wcześniej make sb DO sth – sprawić, że ktoś coś zrobi point OUT sth / point OUT that – wskazać coś / wskazać, że … hear ABOUT sth – usłyszeć o czymś hear FROM sb – dostać wiadomość od kogoś look LIKE + noun – wyglądać jak coś/ktoś come ABOUT (happen) – wydarzyć się; powstać come UP WITH (idea) – wpaść na pomysł set BACK (delay) – opóźnić; zahamować at an ALL-TIME high – na rekordowo wysokim poziomie make / reach / come to a conclusion – wyciągnąć / dojść do wniosku despair – rozpacz; rozpaczać intriguing – intrygujący wearer – osoba nosząca coś sat-nav – nawigacja satelitarna acrobatics – akrobatyka camaraderie – koleżeństwo; poczucie wspólnoty risky / riskiest – ryzykowny / najbardziej ryzykowny chant – skandowanie; przyśpiewka synchronised – zsynchronizowany glamour – blask; splendor compelling – przekonujący; silny maximise – maksymalizować dedicated – przeznaczony wyłącznie do czegoś unsettled – niespokojny; bez stałego miejsca traveller – podróżnik whatsoever – jakikolwiek; w ogóle (w przeczeniach) vibrate – wibrować; drgać vibrations – drgania; wibracje squad – drużyna; zespół coordination – koordynacja paramount – najważniejszy; nadrzędny cheerleading – sportowe układy dopingujące encouragement – zachęta; doping distinctive – charakterystyczny; wyróżniający association – skojarzenie; stowarzyszenie impose – narzucać questionable – wątpliwy rooted in – zakorzeniony w wanderer – wędrowiec lack of dedicated office space – brak przydzielonej przestrzeni biurowej confidante – powiernik / powierniczka toddler – małe dziecko uczące się chodzić reinforce – wzmacniać; utrwalać psychologically – pod względem psychicznym caregiver – opiekun psychological – psychologiczny torment – udręka; dręczyć conclusion – wniosek empowering – dodający sprawczości convincing – przekonujący strive – usilnie się starać regardless – niezależnie od bond – więź variation – odmiana; wariant faintest – najmniejszy (np. faintest idea) incorporate – uwzględniać; włączać straightforward – prosty; nieskomplikowany plausible – wiarygodny; prawdopodobny availability – dostępność armed with – wyposażony w utterance – wypowiedź significant – znaczący setback – niepowodzenie; przeszkoda deliberately – celowo; umyślnie mispronunciation – błędna wymowa linguistically – pod względem językowym acquire – nabyć; przyswoić mistrust – nieufność regret – żałować; żal discomfort – dyskomfort outcome – rezultat persuasive – przekonujący incapable – niezdolny notwithstanding – pomimo (formalnie) mainstay – filar; stały element perform – występować odd-sounding – dziwnie brzmiący refer to – odnosić się do; korzystać z intimidating – onieśmielający viable – wykonalny; realny breadth – szerokość; zakres acquired – nabyty; zdobyty relieve – odciążyć; przynieść ulgę input – wkład; informacje wejściowe insignificant – niezntaczący

Seed Plants

Distance, displacement, speed and displacement

LATE MEDIEVAL AND EARLY RENAISSANCE NORTHERN EUROPE FRAMING THE ERA Rogier van der Weyden and Saint Luke In the 15th century, Flanders—a region corresponding to what is today Belgium, the Netherlands, Luxembourg, and part of northern France (map 20-1)—enjoyed widespread prosperity. Successful merchants and craft guilds joined the clergy and royalty in commissioning artists to produce works for both public and private venues. Especially popular were paintings prepared using the recently per-fected medium of oil-based pigments, which soon became the favored painting medium throughout Europe. One of the early masters of oil painting was Rogier van der Weyden (ca. 1400–1464) of Tournai in present-day Belgium. Rogier made Brussels his home in 1435 and soon thereafter painted Saint Luke Drawing the Virgin (fig. 20-1), probably for the city’s artists’ guild, the Guild of Saint Luke. Luke was the patron saint of artists because legend said he had painted a portrait of the Virgin Mary (see “Early Christian Saints,” pages 246–247). Rogier’s subject was therefore perfectly suited for the head-quarters of a painters’ guild. It shows Luke (his identifying attribute, the ox, is at the right; see “The Four Evangelists,” page 331) at work in the kind of private residence that wealthy Flemish merchants of this era owned. Mary has miraculously appeared before Luke and invited him to paint her portrait as she nurses her son. The saint begins the process by making a preliminary drawing, the same process Rogier himself would have used when he began this commission. The painting thus not only honors Luke but also pays tribute to the profession of painting in Flanders (see “The Artist’s Profession in Flanders,” page 580) by documenting the preparatory work required before artists can begin painting the figures and setting. The subject also draws attention to the venerable history of portrait painting. A rare genre dur-ing the Middle Ages, portraiture became a major source of income for Flemish artists, and Rogier was one of the best portrait painters (fig. 20-12A) in Flanders. In fact, some scholars have suggested that Rogier’s Saint Luke is a self-portrait, identifying the painter with the first Christian artist and under-scoring the holy nature of painting. Saint Luke Drawing the Virgin is also emblematic of 15th-century Flemish painting in aiming to record every detail of a scene with loving fidelity to optical appearance, seen here in the rich fabrics, the patterned floor, and the landscape visible through the window. Also characteristic of Flemish art is the imbuing of many of the painting’s details with symbolic significance. For example, the carved armrest of the Virgin’s bench depicts Adam, Eve, and the serpent, reminding the viewer that Mary is the new Eve and Christ the new Adam who will redeem humanity from Adam and Eve’s Original Sin. 567 Copyright 2021 Cengage Learning. All NORTHERN EUROPE IN THE 15TH CENTURY As the 15th century opened, Rome and Avignon were still the official seats of two competing popes (see “The Great Schism,” page 423), and the Hundred Years’ War (1337–1453) between France and England still raged. The general European movement toward centralized royal governments, begun in the 12th century, continued apace, with a corresponding waning of feudalism. A ris-ing merchant class, whose success depended on individual effort and talent instead of inherited nobility, spawned a new economic system—the early stage of European capitalism. In response to the financial requirements of trade, new credit and exchange systems created a network of enterprising European cities. Trade in money accompanied trade in commodities, and the former financed industry. Both were in the hands of international banking compa-nies, such as those of Jacques Coeur in Bourges (fig. 13-31) and the Medici in Florence (see “The Medici,” page 593). In 1460, Flemish entrepreneurs established the first international commercial stock North Sea E NG L AN D London English Channel Haarlem Ghent Bruges Tournai Amsterdam Nijmegen Louvain Cologne Antwerp Brussels Creglingen Würzburg Paris F R A N C E Tours POITOU Poitiers ATLANTIC OCEAN AUVERGNE Avignon Florence Lucca PAPAL STATES S PA I N 0 0 150 150 300 kilometers Mediterranean Sea 300 miles Rome Tyrrhenian Sea Adriatic Sea OT TOMAN EMP I R E Bourges BERRY Melun Nemours Colmar Dijon Beaune Lake Geneva Geneva Venice Nancy Strasbourg Buxheim H O L Y R O M A N E M P I R E Nuremberg H U NGA RY Kraków Magdeburg PO L AN D exchange in Antwerp. In fact, the French word for stock market (bourse) comes from the name of the van der Beurse family of Bru-ges, the wealthiest city in 15th-century Flanders. Art also thrived in northern Europe during this time, under royal, ducal, church, and private patronage. Two developments in particular were of special significance: the adoption by Rogier van der Weyden (fig. 20-1) and his contemporaries of oil-based pig-ment as the preferred medium for painting, and the blossoming of printmaking as a major art form, which followed Johannes Guten-berg’s invention of moveable type and the printing press. These new media had a dramatic influence on artistic production both north and south of the Alps. THE FRENCH DUCAL COURTS In the 15th century, Flanders was not an independent state but a region under the control of the duke of Burgundy, the ruler of the fertile east-central region of France still famous for its wines (map 20-1). Duke Philip the Bold (r. 1363–1404) was one of four MAP 20-1 France, the duchy of Burgundy, and the Holy Roman Empire in 1477. Late MedievaL and earLy renaissance northern europe 1395–1425 ■ Claus Sluter carves life-size statues of biblical figures with portraitlike features, and Jacques de Baerze and Melchior Broederlam carve and paint the Retable de Champmol for Philip the Bold, duke of Burgundy ■ The Limbourg brothers expand the illusionistic capabilities of manuscript illumination for Jean, duke of Berry 1425–1450 ■ In Flanders, Robert Campin, Jan van Eyck, and Rogier van der Weyden use oil paints to record the exact surface appearance of objects, fabrics, and faces ■ ■ Flemish painters establish portraiture as a major art form German graphic artists pioneer wood-cut printing, making art affordable to the masses 1450–1475 ■ The second generation of Flemish master painters—Petrus Christus, Dieric Bouts, and Hugo van der Goes—continues to use oil paints for altarpieces featuring naturalistic repre-sentations of religious themes ■ In Germany, Johannes Gutenberg invents moveable type and prints the first Bibles on a letterpress 1475–1500 ■ In Flanders, Hans Memling specializes in paintings of the Madonna and Child and portraits of wealthy merchants ■ The Late Gothic style lingers in Ger-many in the large wood altarpieces carved by Veit Stoss and Tilman Riemenschneider ■ Martin Schongauer becomes the first Northern European master of metal engraving sons of King John II (r. 1350–1364) of France. In 1369, Philip mar-ried Margaret de Mâle, the daughter of the count of Flanders, and acquired territory in the Netherlands. Thereafter, the major source of Burgundian wealth was Bruges, the city that made Burgundy a powerful rival of France, which then, as in the Gothic age, was a much smaller kingdom geographically than the modern nation-state. Bruges initially derived its wealth from the wool trade, but soon expanded into banking, becoming the financial clearinghouse for all of northern Europe. Indeed, Bruges so dominated Flanders that the duke of Burgundy eventually chose to make the city his capital and moved his court there from Dijon. Due to the expanded territory and the prosperity of the duchy of Burgundy, Philip the Bold and his successors were probably the most powerful northern European rulers during the first three quarters of the 15th century. Although members of the French royal family, they usually supported England during the Hundred Years’ War because they relied on English raw materials for their wool industry. At times, the dukes of Burgundy controlled much of northern France, including Paris, the seat of the French monarchy. At the height of Burgundian power, the ducal lands stretched from the Rhône River to the North Sea. chartreuse de champmol The great wealth the Burgundy court acquired enabled it to become a significant patron of the arts. Burgundian dukes fully appreciated that artworks could support their dynastic and political goals as well as adorn their castles and townhouses. Philip the Bold’s grandest artistic enterprise was the Chartreuse de Champmol, near Dijon. A chartreuse (“charter house” in English) is a Carthusian monastery. The Carthusian order, founded by Saint Bruno (ca. 1030–1101) in the late 11th century at Chartreuse, near Grenoble in southeastern France, consisted of monks who devoted their lives to solitary liv-ing and prayer. Unlike monastic orders that earned income from farming and other work, the Carthusians generated no revenues. Philip’s generous endowment at Champmol was therefore the sole funding for an ambitious artistic program inspired by the French royal abbey at Saint-Denis (see “Abbot Suger and the Rebuilding of Saint-Denis,” page 383, and fig. 13-1), the burial site of the French kings. The duke’s choice for architect was Drouet de Dammartin, who had worked for his brother King Charles V (r. 1364–1380) on the Louvre (fig. 20-6), the French royal palace in Paris. In 1372, Philip appointed as chief sculptor the Netherlandish master Jean de Marville (d. 1389), who had also worked for Charles V, but at Rouen Cathedral. Emulating the royal burials at Saint-Denis, Philip decided to create at the Dijon chartreuse a ducal mausoleum that would serve both as a means of securing salvation in perpetuity for the ducal family (the monks prayed continuously for their souls) and as a dynastic symbol of Burgundian power. Claus Sluter. In 1389, following the death of Jean de Marville, Philip placed the Neth-erlandish sculptor Claus Sluter (ca. 1360–1406) of Haarlem in charge of the sculptural program (figs. 20-2 and 20-2a). Sluter had joined the Chartreuse de Champmol sculp-ture workshop as Jean’s assistant in 1385. For the portal (fig. 20-2A) of the monastery’s chapel, Sluter’s workshop produced stat-ues of the duke and his wife kneeling before the Virgin and Child. For the cloister, Sluter designed a large sculptural fountain located in a well (fig. 20-2). The well itself served as a water source for the monastery, but Sluter’s fountain did not spout water because the Carthusian commitment to silence and prayer would have precluded anything producing sound. The Well of Moses features statues of Moses and five other Old 20-2a Sluter, Chartreuse de Champmol portal, 1390–1393. 1 ft. 20-2 Claus Sluter, Well of Moses, Chartreuse de Champmol, Dijon, France, 1396–1406. Asnières limestone, painted and gilded by Jean Malouel, Moses 69 high. The Well of Moses, a symbolic fountain of life made for the duke of Bur-gundy, originally supported a sculpted Crucifixion group. Sluter’s figures recall French Gothic jamb statues, but are far more realistic. Testament prophets (David, Daniel, Isaiah, Jeremiah, and Zachariah) ringing a base that once supported a 25-foot-tall group of Christ on the cross mourned by Mary Magdalene, joined perhaps by the Vir-gin Mary and John the Evangelist. Art historians usually describe the Well of Moses as a fons vitae, a fountain of everlasting life, in which the blood of the crucified Christ, symbolized by water, flows down into a basin, and anyone who drinks from the fountain is redeemed. Sluter incorporated The six prophets have almost portraitlike features and distinct individual personalities and costumes. For example, David is an elegantly garbed Gothic king and Moses an elderly horned prophet (compare fig. 12-38) with a waist-length beard. Sluter intensely studied the natural appearance of faces and fabrics in order to sculpt the biblical figures in minute detail. Heavy draperies with voluminous folds swathe the life-size statues. Sluter succeeded in making their difficult, complex surfaces seem remarkably natural-istic. He enhanced this effect by skillfully differentiating textures, from coarse drapery to smooth flesh and silky hair. Paint, much of which has flaked off, further augmented the naturalism of the figures. (The painter was Jean Malouel [ca. 1365–1415], another Netherlandish master.) This fascination with the specific and tan-gible in the visible world became one of the chief characteristics of 15th-century Flemish art. Retable de Champmol. Philip the Bold also commissioned a major altarpiece—the Retable de Champmol (figs. 20-3 and 20-4)—for the main altar in the chapel of the Chartreuse. A collaborative project between two Flemish artists, this altarpiece consisted of a 1 ft. 20-3 Jacques de Baerze, Retable de Champmol (open), from the chapel of the Chartreuse de Champmol, Dijon, France, installed 1399. Wood, center section 59 5 3 40 × 89 3 1 40. Musée des Beaux-Arts, Dijon. Worshipers could view the interior of the Champmol altarpiece on Sundays and feast days. Jacques de Baerze carved three scenes of Jesus’s life centered on the Crucifixion, a fitting backdrop for the Eucharist. 20-4 Melchior Broederlam, Retable de Champmol (closed), from the chapel of the Chartreuse de Champmol, Dijon, France, installed 1399. Oil and tempera on wood, each wing 59 5 3 Arts, Dijon. 40 × 49 1 1 40. Musée des Beaux-This early example of oil painting attempts to represent the three-dimensional world on a two-dimensional surface, but the gold background and flat halos recall medieval pictorial conventions. 1 the symbolism of the fons vitae in his waterless fountain, in which the viewer must imagine Christ’s blood stream-ing down over grieving angels and the Savior’s prophetic predeces-sors. Although the models for the Old Testament figures may have been French Gothic cathedral portal statues, as was certainly the case for the Dijon chapel statues (fig. 20-2A), the inspiration for the Well of Moses prophets may have come more directly from contempora-neous mystery plays in which actors portraying prophets frequently delivered commentaries on events in Christ’s life. large sculptured shrine (fig. 20-3) by Jacques de Baerze (active ca. 1384–1399) and a pair of painted exterior panels (fig. 20-4) by Melchior Broederlam (active ca. 1387–1409). Altarpieces were a major art form north of the Alps during the late 14th and the 15th century. From their position behind the altar, they served as backdrops for the Mass, the ritual celebration of the Holy Eucharist. At the Last Supper, Christ commanded his apostles to reenact the Eucharistic meal and repeat in memory of him the communion credo, wherein he tenders his body to eat and his blood to drink (see “The Life of Jesus in Art,” pages 244–245). This act serves as the nucleus of the Mass, which involves this reenactment as well as prayer and contemplation of the word of God. Because the Mass involves not only a memorial rite but Christian doctrine as well, art has traditionally played an important role in giving visual form to complex theological concepts for the faithful. Like the narrative reliefs in medieval church portals, these altarpieces had an important teaching function, especially for the illiterate. By reinforcing Church doctrines visually, they also stimulated devotion. These altarpieces most often took the form of polyptychs—hinged multipaneled paintings or multiple carved relief panels. The hinges enabled the clergy to close the polyptych’s side wings over the central panel(s). Because artists decorated both the exterior and inte-rior of the altarpieces, this multi-image format provided the opportu-nity to construct narratives through a sequence of images. Although concrete information is lacking about when the clergy opened and closed the altarpieces, the wings probably normally remained closed but were opened on Sundays and feast days. On this schedule, view-ers could have seen both the interior and exterior—diverse imagery at various times according to the liturgical calendar. Given their placement in churches as backdrops to the Mass, it is not surprising that among the many kinds of scenes depicted on altarpieces, themes directly related to Christ’s sacrifice are com-mon. The Champmol altarpiece, or retable, for example, features sculpted passion scenes on the interior: the Crucifixion (at the cen-ter) and (at left and right respectively) the adoration of the magi and the entombment of Christ, with rows of saints on the wings, all framed by delicate pinnacles, tracery, and lancets. The carved wood narratives span the full life cycle of Jesus from his birth to his death and burial, with the figure of the Savior on the cross forming a fit-ting backdrop to the Eucharist. On the Champmol retable’s painted wings (fig. 20-4), Broeder-lam depicted the Annunciation and Visitation (on the left panel) and the presentation in the temple and flight into Egypt (on the right). Thus Broederlam’s painted images on the altarpiece’s exterior gave a fuller account of the Savior’s early life and set the stage for de Baerze’s passion scenes when the wings were folded back to reveal the sculpted narratives. The two painted wings include both landscape and inte-rior scenes. Broederlam depicted the buildings in both Romanesque and Gothic styles. Scholars have suggested that the juxtaposition of different architectural styles in the left panel is symbolic. The rotunda (round building, usually with a dome, a common antique type) refers to the Old Testament, whereas the “modern” Gothic porch with its lancet windows and tracery relates to the New Testament. In the right panel, a statue of a Greco-Roman god falls from the top of a column as the holy family approaches. These and other details announce the coming of the new order under Christ. Stylistically, Broederlam’s panels are a mixture of three-dimen-sional rendition of the landscape and buildings with a solid gold background and flat gold haloes for the holy figures, regardless of the positions of their heads. Despite these lingering medieval pictorial conventions, the altarpiece is an early example of many of the artistic developments that preoccupied European artists throughout the 15th century, especially the illusionistic depiction of three-dimensional objects and the naturalistic representation of landscape. Oil Painting. The Retable de Champmol also foreshadowed an-other significant development in 15th-century art: the widespread adoption of oil paints, which Broederlam mixed with tempera in this early example (see “Tempera and Oil Painting,” page 574). Oil paints facilitated the exactitude in rendering details so charac-teristic of Northern European painting. Although the Italian biogra-pher Giorgio Vasari (see “Giorgio Vasari’s Lives,” page 648) and other 16th-century commentators credited the Flemish master Jan van Eyck (discussed later in this chapter) with the invention of oil paint-ing, recent evidence has revealed that oil paints had been known for some time, well before Melchior Broederlam liberally used oils for Philip the Bold’s Dijon altarpiece. Flemish painters built up their pic-tures by superimposing translucent paint layers on a layer of under-painting, which in turn had been built up from a carefully planned drawing (compare Saint Luke at work in fig. 20-1) made on a panel prepared with a white ground. With the oil medium, artists could create richer colors and more subtle gradations of tone than previ-ously possible, giving their paintings the illusion of glowing light and glistening surfaces. These traits differed significantly from the high-keyed color, sharp light, and rather matte (dull) surface of tempera. The brilliant and versatile oil medium suited perfectly the formal intentions of the generation of Flemish painters after Broederlam, who aimed for sharply focused clarity of detail in their representation of objects ranging in scale from large to almost invisible. Manuscript painting During the 15th century, Northern European artists built on the achievements of Gothic painters (see “Gothic Book Production,” page 402) and produced exquisitely refined illuminated manu-scripts. Among the most significant developments in manuscript painting was a new interest in representing fully modeled figures moving freely in convincing three-dimensional settings. Increased contact with Italy, where Renaissance artists had revived the picto-rial principles of classical antiquity, may have influenced Northern painters’ interest in illusionism. Limbourg Brothers. The most innovative early-15th-century manuscript illuminators were the three Limbourg brothers—Pol, Herman, and Jean—from Nijmegen in the Netherlands. They were nephews of Jean Malouel, the court artist of Philip the Bold, who painted Sluter’s Well of Moses statues (fig. 20-2). Following in the footsteps of earlier illustrators such as Jean Pucelle (figs. 13-36 and 13-36A), the Limbourg brothers expanded the illusionistic capabili-ties of illumination. Trained in the Netherlands, the brothers moved to Paris no later than 1402, and between 1405 and their death in 1416 (probably from the plague), they worked in Paris and Bourges for Jean, duke of Berry (r. 1360–1416) and brother of Philip the Bold of Burgundy and King Charles V (r. 1364–1380) of France. Jean ruled the western French regions of Berry, Poitou, and Auvergne (map 20-1). The duke was an avid art patron and focused on collecting manuscripts, jewels, and rare artifacts. Among the more than 300 manuscripts he owned were Pucelle’s Belleville Breviary (fig. 13-36) and the Hours ofJeanne d’Evreux (fig. 13-36A) as well as Les Très Riches Heures du Duc de Berry (The Very Sumptuous Hours of the Duke of Berry; figs. 20-5 and 20-6), which he commissioned the Limbourg brothers to produce. A Book of Hours, like a breviary, was The French Ducal Courts Limbourg brothers (Pol, Herman, Jean), January, from Les Très Riches Heures du Duc de Berry, 1413–1416. Colors and ink on vellum, 8 7 80 × 5 3 80. Musée Condé, Chantilly. The pictures in Les Très Riches Heures depict characteristic activities of each month. The prominence of genre subjects reflects the increasing inte-gration of religious and secular art in the 15th century. a book used for reciting prayers (see “Medieval Books,” page 324). As prayer books, they replaced the traditional psalters (books of psalms), which were the only liturgical books in private hands until the mid-13th century. Books of Hours contained liturgical passages to be read privately at set times during the day, from matins (dawn prayers) to compline (the last of the prayers recited daily), including the “Office [prayer] of the Blessed Virgin.” An illustrated calendar containing local religious feast days usually preceded the Office of the Blessed Virgin. Penitential psalms, devotional prayers, litanies to the saints, and other prayers, including those of the Holy Cross and for the dead, followed. Books of Hours became favorite possessions of the Northern European aristocracy during the 14th and 15th centuries. These costly books eventually became available to affluent merchants and contributed to the decentralization of religious practice that was one factor in the Protestant Reformation in the early 16th century (see “Luther and the Reformation,” page 692). The full-page calendar pictures of Les Très Riches Heures are the most famous in the history of manuscript illumination. They 20-6 Limbourg brothers (Pol, Herman, Jean), October, from Les Très Riches Heures du Duc de Berry, 1413–1416. Colors and ink on vellum, 8 7 80 × 5 3 80. Musée Condé, Chantilly. The Limbourg brothers were pioneers in expanding the illusionistic capabili-ties of manuscript painting with their care in rendering architectural details and their convincing depiction of cast shadows. represent the 12 months in terms of the associated seasonal tasks, alternating scenes of nobility and peasantry, and featuring the duke’s relationship with his courtiers and peasants. Above each picture is a lunette in which the Limbourgs depicted the zodiac signs and the chariot of the sun as it makes its yearly cycle through the heavens. Beyond its function as a religious book, Les Très Riches Heures also furnishes a picture of life in the territory the duke ruled—a picture designed to flatter the Limbourgs’ patron. For example, the color-ful calendar picture for January (fig. 20-5) portrays a New Year’s reception at court. The duke appears as magnanimous host, his head circled by the fire screen, almost halolike, behind him. His chamberlain stands next to him, urging the guests forward with the words “aproche, aproche,” words that call attention to the Lim-bourgs’ success in creating the illusion of deep spatial recession. The lavish spread of food on the table and the large tapestry on the back wall augment the richness of the setting and the occasion By contrast, the illustration for October (fig. 20-6) focuses on the peasantry. Here, the Limbourg brothers depicted a sower, a man plowing on horseback, and washerwomen, along with city dwell-ers, who promenade in front of the Louvre, the French king’s resi-dence at the time, now one of the world’s great art museums. (Paris was not then the highly populated urban center it is today. Open fields surrounded the royal palace.) The peasants do not appear dis-contented as they go about their various tasks. Surely this imagery flattered the duke’s sense of himself as a compassionate master. The growing artistic interest in naturalism is evident here in the careful way the painter recorded the architectural details of the Louvre and in the convincing shadows of the people and objects (such as the archer scarecrow and the horse) in the scene. As a whole, Les Très Riches Heures reinforced the image of the duke of Berry as a devout man, cultured bibliophile (“lover of books”), sophisticated art patron, and powerful and magnanimous leader. Further, the expanded range of subject matter, especially the prominence of genre subjects in a religious book, reflected the increasing integration of religious and secular concerns in both art and life during the 15th century. Although all three Limbourg brothers worked on Les Très Riches Heures, art historians have never been able to ascertain definitively which brother painted which images. (One scholar has suggested that because the book was left unfinished at the time of the brothers’ deaths, some of the details are the work of a fourth artist, who completed the illustrations around the middle of the century.) Given the common practice of collaboration on artistic projects at this time, however, the determination of specific author-ship is not very important. Together, the Très Riches Heures illumi-nators reoriented the art of manuscript illumination to approximate more closely the art of panel painting, treating the pages of books as if they were windows onto the world of 15th-century Europe. Mary of Burgundy. The Limbourgs’ most accomplished suc-cessor at the end of the century was an anonymous illuminator known as the Master of Mary of Burgundy. Some scholars identify him as Alexander Bening (ca. 1444–1519) of Ghent, a painter in the circle of Hugo van der Goes (fig. 20-16). The two facing pages illustrated here (fig. 20-7) are the opening full-page illumination and one of the daily prayers in the private devotional Book of Hours that belonged to Mary, the last duchess of Burgundy (r. 1477–1482) and daughter of Charles the Bold. She is the elegantly dressed woman sitting in a private chapel with her pet dog on her lap, reading from a prayer book of the same size and format as the one in which her portrait appears. On a ledge to her left are her rosary beads and a tall vase with purple irises—flowers associated tempera and oil painting The generic words paint and pigment encompass a wide range of substances that artists have used through the ages. Fresco aside (see “Fresco Painting,” page 428), during the 14th century, egg tempera was the material of choice for most painters, both in Italy and northern Europe. Tempera consists of egg combined with a wet paste of ground pigment. In his influential 1437 guidebook Il libro dell’arte (The Handbook of Art; see “Imitation and Emulation in Renaissance Art,” page 606), Cennino Cennini (ca. 1370–ca. 1440) noted that artists mixed only the egg yolk with the ground pigment, but analyses of paintings from this period have revealed that some artists chose to use the entire egg. Images painted with tempera have a velvety sheen. Artists usually applied tempera to the painting surface with a light touch because thick application of the pig-ment mixture would result in premature cracking and flaking. Some artists used oil paints (powdered pigments mixed with lin-seed oil) as far back as the eighth century, but not until the early 1400s did oil painting become widespread. Melchior Broederlam (fig. 20-4), Robert Campin (fig. 20-8), and other Flemish artists were among the first to employ oils extensively, often mixing them with tempera, as Broederlam did. (Italian painters soon followed suit, underscoring that artistic exchanges across the Alps went in both directions during the Renaissance.) The discovery of better drying components in the early 15th century enhanced the setting capabilities of oils. Rather than apply these oils in the light, flecked brushstrokes that the tempera technique encouraged, artists laid down the oils in transparent layers, or glazes, over opaque or semiopaque underlayers. In this manner, painters could build up deep and subtly gradated tones through repeated glazing. Unlike works in tempera, whose surface dries quickly due to water evaporation, oils dry more uniformly and slowly, giving the artist time to rework areas. This flexibility must have been particularly appealing to artists who worked very deliberately, such as Campin, Jan van Eyck, Rogier van der Wey-den, and the other Flemish masters discussed in this chapter, as well as the Italian Leonardo da Vinci (figS. 22-2 and 22-5). Leonardo also preferred oil paint because its gradual drying process and consistency enabled him to blend the pigments, thereby creating the impressive sfu-mato (smoky) effect that contributed to his fame. Moreover, while dry-ing, oil paints smooth out, erasing any trace of the brush that applied the paint. Oil paints also produce a glowing surface, creating a rich visual effect unlike the duller sheen of tempera. Both tempera and oils can be applied to various surfaces. Through the early 16th century, wood panels served as the foundation for most paintings. Italians painted on poplar. Northern European artists used oak, lime, beech, chestnut, cherry, pine, and silver fir. Local availability of these timbers determined the choice of wood. Linen canvas became increasingly popular in the late 16th century. Although evidence sug-gests that artists did not intend permanency for their early images on canvas, the material proved particularly useful in areas such as Venice where high humidity warped wood panels and made fresco unfeasible. Furthermore, until artists began to use wood bars to stretch the can-vas to form a taut surface, canvas paintings could be rolled and were lighter and more compact and therefore more easily portable than wood panels. with the Virgin Mary. The masterful representation of rich fabrics, Mary’s transparent veil, and the still life of flowers all attest to this painter’s skill and powers of observation. Even more remarkable is the illuminator’s meticulously detailed depiction of the Gothic church seen through the window of Mary’s chapel, especially because the manuscript page is only half the size of Jan van Eyck’s Madonna in a Church (fig. 20-10A). The scene presented to the viewer through the chapel window within the page’s “window onto the world”—a brilliant conception in itself—is the realization of Mary’s prayers. The duchess has been granted a private audience with the Virgin without any interced-ing priest. She, accompanied by her ladies-in-waiting, kneels before the Virgin, who sits in front of the church altar, attended by angels and holding the Christ Child in her lap. Both holy figures turn their heads to look at the duchess. The illumination has few peers in the history of art as an image of pious devotion. FLANDERS As noted, the greatest single source of income for the Burgundian ducal court came from banking and trade activities in Flanders, ter-ritory that Philip the Bold acquired when he married the Flemish count’s daughter. Flanders was also the leading artistic center of Northern Europe in the 15th century. Master of Flémalle One of the greatest Flemish painters of the generation after Melchior Broederlam was the man known as the Master of Flémalle, whom most art historians identify as Robert Campin (ca. 1375–1444), the leading painter of Tournai, who also worked in the newly popular medium of oil painting (see “Tempera and Oil Painting,” page 574). His most famous work is the Mérode Triptych (fig. 20-8), although it has recently been suggested that the triptych (three-panel painting) is the work of more than one artist and that the wings are a later addition. The question of authorship apart, the painting is one of the many small works of this period produced for private patrons and intended for household prayer. Perhaps the most striking feature of these private devotional images is the integration of religious and secular concerns. For example, artists often presented biblical scenes as taking place in a house (compare fig. 20-1). Religion was such an integral part of Flemish life that separating the sacred from the secular was almost impossible—and undesirable. Furthermore, the presentation in religious art of famil-iar settings and objects no doubt strengthened the direct bond that the patron or viewer felt with biblical figures. The Annunciation, prophesied in Isaiah 7:14, is the subject of the central panel of Campin’s triptych. Unseen by the Virgin, a tiny figure of Christ carrying the cross of his martyrdom and of human salvation enters the room on a ray of light, a potent symbol of the Incarnation and a foreshadowing of what lies ahead for the future Mother of God and her son. The archangel Gabriel approaches Mary, who sits reading inside a well-kept Flemish home. Through the window in the background of the right wing, the viewer sees a local cityscape, as in Rogier van der Weyden’s later Saint Luke (fig. 20-1). The depicted accessories, furniture, and utensils con-firm the locale as Flanders. However, the objects represented are not merely decorative. They also function as religious symbols. The book, extinguished candle, and lilies on the table; the copper basin in the corner niche; the towels, fire screen, and bench all symbolize, in different ways, the Virgin’s purity and her divine mission. In the right panel, Joseph, apparently unaware of the angel’s arrival, has constructed two mousetraps, symbolic of the theologi-cal concept that Christ is bait set in the trap of the world to catch the Devil. The ax, saw, and rod that Campin painted in the fore-ground of Joseph’s workshop not only are tools of the carpenter’s trade but also are mentioned in Isaiah 10:15. In the left panel, the closed garden is symbolic of Mary’s purity, and the flowers Campin included relate to Mary’s virtues, especially humility. The triptych’s donor, Peter Inghelbrecht, a wealthy merchant, and his wife, Margarete Scrynmakers, kneel in the garden and witness the momentous event through an open door. Donor portraits—portraits of the individual(s) who commissioned (or “donated”) the work—became very popular in the 15th century. In this instance, in addi-tion to asking to be represented in their altarpiece, the Inghelbrechts probably specified the subject. Inghelbrecht means “angel bringer,” a reference to Gabriel’s bringing the news of Christ’s birth to Mary in the central panel. Scrynmakers means “cabinet-or shrine-makers,” and probably inspired the workshop scene in the right panel. Jan van eyck In 1425, Jan van Eyck (ca. 1395–1441), the most famous of several internationally renowned Flemish artists, became the court painter of Philip the Good, duke of Burgundy (r. 1419–1467). In 1431, he moved his studio to Bruges, where the duke maintained his official residence. Ghent Altarpiece. The year Jan arrived in Bruges, he set out to complete the Ghent Altarpiece (figs. 20-9 and 20-10), which, according to the dedicatory inscription, his older brother, Hubert van Eyck (ca. 1385–1426), had begun. Given Jan’s stature today, it is noteworthy that the inscription describes Hubert as an artist “than whom there was no greater” and Jan as “his brother, second in art.” The nearly 12-foot-tall Ghent retable is one of the largest of the 15th century. Jodocus Vyd (d. 1439), diplomat-retainer of Philip the Good, and his wife, Elisabeth Borluut (d. 1443), commissioned the polyptych as the centerpiece of the chapel Vyd built in the Ghent church originally dedicated to Saint John the Baptist (since 1540, Saint Bavo Cathedral). Vyd’s largesse and the political and social connections that the Ghent Altarpiece revealed to its audience contributed to Vyd’s appointment as the city’s burgomeister (chief magistrate; mayor) shortly after the unveiling of the work. Two of the exterior panels (fig. 20-9) depict the donors. The husband and wife, in paired niches with Gothic tracery, kneel with their hands clasped in prayer. They gaze piously at two illusionistically rendered statues (painted in grisaille—that is, solely in gray tones to imitate stone) that reflect the innovative style of Claus Sluter (figs. 20-2 and 20-2A). The sculptures represent Ghent’s patron saint, John the Baptist, and John the Evangelist, who was probably Vyd’s patron saint. The Annunciation appears on the upper register, with a care-ful representation of a Flemish town outside the painted window of the center panel (compare figs. 20-1 and 20-8). In the uppermost arched panels, Jan depicted the Old Testament prophets Zachariah and Micah, along with sibyls, Greco-Roman mythological female prophets whose writings the Church interpreted as prophecies of Christ. An interesting detail is the exchange of words between Gabriel and the Virgin. Jan painted the words of Mary’s response upside down, because they are directed to God in Heaven. When open (fig. 20-10), the altarpiece reveals a richly colored painting of human redemption through Christ. In the upper regis-ter, God the Father—wearing the pope’s triple tiara, with a worldly Flanders 575 Copyright 20-9 Hubert and Jan van Eyck, Ghent Altar-piece (closed), Saint Bavo Cathedral, Ghent, Belgium, ca. 1423–1432. Oil on wood, 119 50 × 79 60. Grandiose multipanel painted altarpieces were popular in Flemish churches. Artists decorated both the interiors and exteriors of these polyp-tychs, which often, as here, included donor portraits. 1 ft. 1 ft. 20-10 Hubert and Jan van Eyck, Ghent Altarpiece (open), Saint Bavo Cathedral, Ghent, Belgium, ca. 1423–1432. Oil on wood, 119 50 × 159 10. In this richly colored painting of salvation from the Original Sin of Adam and Eve, God the Father presides in majesty. The van Eyck brothers used oil paints to render every detail with loving fidelity to appearance. 576 crown at his feet and resplendent in a deep-scarlet mantle—presides in majesty. To God’s right is the Virgin, represented, as in the Gothic age and in a small Jan van Eyck diptych (two-paneled painting; fig. 20-10A), as the queen of Heaven, with a crown of 12 stars on her head. John the Baptist sits to God’s left. To either side is a choir of angels, with an angel playing an organ on the right. Adam and Eve appear in the far panels. The inscriptions in the arches above Mary and Saint John extol the Virgin’s virtue and purity and Saint John’s greatness as the forerunner of Christ (see “Early Christian Saints,” pages 246–247). The inscription above the Lord’s head translates as “This is God, all-powerful in his divine majesty; of all the best, by the gentle-ness of his goodness; the most liberal giver, because of his infinite a second opinion Giovanni Arnolfini and His Wife In 1434, Jan van Eyck depicted Giovanni Arnolfini, a Lucca financier who had established himself in Bruges as an agent of the Medici family, and his second wife in their home (fig. 20-11). Arnolfini holds the hand of his spouse, whose name is not known. That much is certain, but the purpose and meaning of the double portrait remain the subject of con-siderable debate, with a few new interpretations having been advanced just during the last several years. According to the traditional interpreta-tion of the painting, Jan recorded the couple taking their marriage vows. As in the Mérode Triptych (fig. 20-8), almost every object portrayed car-ries meaning. For example, the little dog symbolizes fidelity (the com-mon canine name Fido originated from the Latin fidere, “to trust”). The finial (crowning ornament) of the marriage bed at the right is a tiny statue of Saint Margaret, patron saint of childbirth. (The bride is not yet preg-nant, although the fashionable costume she wears makes her appear so.) From the finial hangs a whiskbroom, symbolic of domestic care. Indeed, even the placement of the two figures in the room is meaningful. The woman stands near the bed and well into the room, whereas the man stands near the open window, symbolic of the outside world. Many art historians, however, dispute this interpretation because, among other things, the room in which Arnolfini and his wife stand is a public reception area, not a bedchamber. One scholar has suggested that Arnolfini is conferring legal privileges on his wife to conduct busi-ness in his absence. Another thinks that this may be a memorial portrait of a wife who died in childbirth. In any case, an important aspect of the painting is that the artist functions as a witness to whatever event is taking place. In the back-ground, framed by the arms and joined hands of the two figures, is a convex mirror (complete with its spatial distortion, brilliantly recorded; compare fig. 22-40), in which Jan depicted not only the principals, Arnolfini and his wife, but also two persons who look into the room through the door. (Arnolfini’s raised right hand may be a gesture of greeting to the two men.) One of these must be the artist himself, as the elegant inscription above the mirror, Johannes de Eyck fuit hic (“Jan van Eyck was here”), announces that he was present. The self-portrait also underscores the painter’s self-consciousness as a professional art-ist whose role deserves to be recorded and remembered. generosity.” The step behind the crown at the Lord’s feet bears the inscription “On his head, life without death. On his brow, youth without age. On his right, joy without sadness. On his left, security without fear.” The entire altarpiece amplifies the central theme of salvation. Even though humans, personified by Adam and Eve, are sinful, they will be saved because God, in his infinite love, will sac-rifice his own son for their sake. The panels of the lower register extend the symbolism of the upper. In the center panel, saints arrive from the four corners of the earth through an opulent, flower-spangled landscape. They pro-ceed toward the altar of the Lamb and the octagonal fountain of life (compare fig. 20-2). The book of Revelation passage that recounts the adoration of the Lamb is the main reading on All Saints’ Day (November 1). The Lamb symbolizes the sacrificed son of God, whose heart bleeds into a chalice, while into the fountain spills the “pure river of water of life, clear as crystal, proceeding out of the throne of God and of the Lamb” (Rev. 22:1). On the right, the 1 ft. 20-11 Jan van Eyck, Giovanni Arnolfini and His Wife, 1434. Oil on wood, 29 90 × 19 10 1 20. National Gallery, London. Jan van Eyck played a major role in establishing portraiture as an important Flemish art form. In this portrait of an Italian financier and his wife, he also portrayed himself in the convex mirror on the rear wall. Flanders Framed paintings Until the mid-20th century, when painters began to affix canvas to simple wood stretcher bars to provide a taut painting surface devoid of orna-mentation, artists considered the frame an integral part of the painting. Frames served a number of functions, some visual, others conceptual. For paintings such as large-scale altarpieces that were part of a larger environment, frames often served to integrate the painting with its sur-roundings. Frames could also be used to reinforce the illusionistic nature of the painted image. For example, the Italian painter Giovanni Bellini, in his San Zaccaria Altarpiece (fig. 22-29), duplicated the carved pilasters of the architectural frame in the painting itself, thereby enhancing the illusion of space and giving the painted figures an enhanced physical presence. In the Ghent Altarpiece, the frame seems to cast shadows on the floor between the angel and Mary in the Annunciation scene (fig. 20-9, top). More commonly, artists used frames specifically to dis-tance the viewer from the (often otherworldly) scene by calling attention to the separation of the image from the viewer’s space. Most 15th-and 16th-century paintings included elaborate frames that the artists themselves helped design and construct. Extant con-tracts reveal that the frame could account for as much as half of the cost of an altarpiece. Frequently, the commissions called for painted or gilded frames, adding to the expense. For small works, artists some-times affixed the frames to the panels before painting, so that the frame was a point of reference as they worked. Occasionally, a single piece of wood served as both panel and frame, and the artist (or, in practice, an assistant) carved the painting surface from the wood, leaving the edges as a frame. Larger images with elaborate frames, such as altarpieces, required the services of a professional woodcarver or stonemason. The painter worked closely with the individual constructing the frame to ensure its appropriateness for the image(s) produced. Unfortunately, over time, many frames have been removed from their paintings. For instance, in 1566 church officials dismantled the Ghent Altarpiece and detached its elaborately carved frame in order to protect the sacred work from Protestant iconoclasts (see “Art and the Reformation,” page 692). As ill luck would have it, when the panels were reinstalled in 1587, no one could find the frame. Sadly, the absence of many of the original frames of old paintings deprives viewers today of the painter’s complete artistic vision. Conversely, when the original frames exist, they sometimes provide essential information, such as the 1 in. 20-12 Jan van Eyck, Man in a Red Turban, 1433. Oil on wood, 19 11 80 × 10 1 40. National Gallery, London. Man in a Red Turban is the first known Western painted portrait in a thousand years in which the sitter looks directly at the viewer. The inscribed frame suggests that it is a self-portrait of Jan van Eyck. subject, name of the painter, and date. For example, the inscriptions on the frame of Jan van Eyck’s Man in a Red Turban (fig. 20-12) state that he painted it on October 21, 1433, and the inclusion of “As I can” and omission of the sitter’s name suggest that the painting is a self-portrait. 12 apostles and a group of martyrs in red robes advance. On the left appear prophets. In the right background come the virgin martyrs, and in the left background the holy confessors approach. On the lower wings, hermits, pilgrims, knights, and judges enter from left and right. They symbolize the four cardinal virtues: Temperance, Prudence, Fortitude, and Justice, respectively. The altarpiece cele-brates the whole Christian cycle from the fall of man to the redemp-tion, presenting the Church triumphant in heavenly Jerusalem. Jan used oil paints to render the entire altarpiece in shim-mering colors that defy reproduction. No small detail escaped the painter. With pristine specificity, he revealed the beauty of the most insignificant object. He depicted the soft texture of hair, the glitter of gold in the heavy brocades, the luster of pearls, and the flash-ing of gems, all with loving fidelity to appearance. This kind of meticulous attention to recording details became a hallmark of Flemish panel painting in the 15th century. Giovanni Arnolfini. Emerging capitalism led to prosperity in Europe’s booming urban centers and fueled in turn the growing bourgeois market for art objects, particularly in Bruges, Antwerp, and, later, Amsterdam. The increased wealth of the merchant class contributed to an expanded market for secular art in addition to religious artworks. It is noteworthy, for example, that both the Mérode Triptych and the Ghent Altarpiece—certainly to be catego-rized as “religious art”—include painted portraits of their donors. These paintings marked a significant revival of portraiture, a genre that had languished since antiquity. Private commissions for por-traits as independent artworks began to multiply as both artists and 578 patrons became interested in the reality (both physical and psycho-logical) that portraits could reveal. In the 15th century, Flemish patrons eagerly embraced the opportunity to have their likenesses painted. The elite wanted to memorialize themselves in their dynastic lines and to establish their identities, ranks, and stations with images far more concrete than heraldic coats of arms. Portraits also served to represent state offi-cials at events they could not attend. Royalty, nobility, and the very rich would sometimes send artists to paint the likeness of a pro-spective bride or groom. For example, when young King Charles VI (r. 1380–1422) of France sought a bride, he dispatched a painter to three different royal courts to make portraits of the candidates. But prosperous merchants also commissioned portraits for their homes. An early example of secular portraiture is Jan van Eyck’s dif-ficult-to-interpret depiction of the banker Giovanni Arnolfini (see “Giovanni Arnolfini and His Wife,” page 577). Man in a Red Turban. Whatever the intended meaning of Giovanni Arnolfini and His Wife, the painting is representative of the growing importance of secular portraiture as an income-producing artistic genre in 15th-century Flanders. In Man in a Red Turban (fig. 20-12), the man whom Jan van Eyck portrayed looks directly at the viewer. This is the first known Western painted portrait in a thousand years where the sitter does so. The level, composed gaze, directed from a true three-quarter head pose, must have impressed observers deeply. The painter created the illu-sion that from whatever angle a viewer observes the face, the eyes return that gaze. Jan, with his considerable observational skill and controlled painting style, injected a heightened sense of specificity into this portrait by including beard stubble, veins in the bloodshot left eye, and weathered, aged skin. Although a definitive identifica-tion of the sitter has yet to be made, most art historians consider Man in a Red Turban a self-portrait, which Jan painted by looking at his image in a mirror (as he depicted himself in the mirror in the Arnolfinis’ home; fig. 20-11). The inscriptions on the frame (see “Framed Paintings,” page 578) reinforce this identification. Across the top, Jan wrote “As I can” in Flemish using Greek letters. One suggestion is that this portrait was a demonstration piece intended for prospective clients, who could compare the painting with the painter and judge what he “could do” in terms of recording a faith-ful likeness. Across the bottom appear the date and a statement in Latin: “Jan van Eyck made me.” The use of both Greek and Latin suggests that the artist viewed himself as both a learned man and a worthy successor to the fabled painters of antiquity. rogier van der Weyden When Jan van Eyck began work on the Ghent Altarpiece, Rogier van der Weyden (fig. 20-1) was an assistant in the work-shop of Robert Campin (fig. 20-8), but the younger Tournai painter’s fame soon eclipsed Campin’s and eventually rivaled Jan’s. Rogier’s renown rested on his skill in recording indi-vidual features and character in his portraits (fig. 20-12a) and especially on the dynamic compositions of his narrative works, which stress human action and drama. He concen-trated on Christian themes, especially those episodes in the life of Jesus that elicited pow-erful emotions—for example, the Crucifixion and Pietà (the Vir-gin Mary cradling the dead body of her son)—moving observers deeply by vividly portraying the sufferings of Christ. 20-12a Van der Weyden, Portrait of a Lady, ca. 1460. Deposition. One of Rogier’s early masterworks is Deposition (fig. 20-13), the center panel of a triptych commissioned by the archers’ guild of Louvain for the church of Notre-Dame hors-les-murs (Church of Our Lady—the Virgin—outside the [town] walls). Rogier acknowledged the patrons of this large painting by incorpo-rating the crossbow (the guild’s symbol) into the decorative trac-ery in the corners. Instead of creating a deep landscape setting, as Jan van Eyck might have, Rogier compressed the figures and action onto a shallow stage with a golden back wall, imitat-ing the large sculptured shrines so popular in the 15th century, of which Jacques de Baerze’s Retable de Champ-mol (fig. 20-3) is one of many Nether-landish examples (compare figs. 20-21 and 20-22). The device admirably served his purpose of expressing max-imum action within a limited space 20-13 Rogier van der Weyden, Deposition, center panel of a triptych from Notre-Dame hors-les-murs, Louvain, Belgium, ca. 1435–1442. Oil on wood, 79 2 5 del Prado, Madrid. 80 × 89 7 1 1 ft. 80. Museo Deposition resembles a relief carving in which the biblical figures act out a drama of passionate sorrow as if on a shallow theatrical stage. The painting makes an unforgettable emotional impression. and dramatically confronting the viewer, but the setting of the Cru-cifixion in a box is unrealistic, as is the size of the cross, the arms of which are not wide enough for Jesus’s hands to have been nailed to them. The painting, with the artist’s crisp drawing and precise mod-eling of forms, resembles a stratified relief carving, and the viewer may wonder if this is a painting of the biblical event or a painting of a shrine representing the event. In any case, a series of lateral undu-lating movements gives the group a compositional unity, a formal cohesion that Rogier strengthened by depicting the sorrowful anguish that many of the figures share. Present are the Virgin, sev-eral of her half-sisters, Joseph of Arimathea, Nicodemus, Saint John the Evangelist, and Mary Magdalene. The similar poses of Christ and his mother further unify the composition, evoke the shape of the crossbows in the corner tracery, and reflect the belief that Mary suffered the same pain at the Crucifixion as her son. ART AND SOCIETY the artist’s profession in Flanders As they did in Italy (see “Artistic Training,” page 434), guilds controlled artistic production in Flanders. To pursue a craft, individuals had to belong to the guild representing that craft. Painters, for example, sought admission to the Guild of Saint Luke, the patron saint of painters because Luke made a portrait of the Virgin Mary (fig. 20-1). The patron saint of the metalworkers’ guild was Saint Eligius (fig. 20-14), who was a goldsmith before devoting himself to the Church and eventually becom-ing bishop of Tournai. The path to eventual membership in the guild began, for men, at an early age, when the father apprenticed his son in boyhood to a mas-ter, with whom the young aspiring painter lived. The master taught the fundamentals of his craft—how to make implements, prepare panels with gesso (plaster mixed with a binding material), and mix colors, oils, and varnishes. Once the youth mastered these procedures and learned to work in the master’s traditional manner, he usually spent several years working as a journeyman in various cities, observing and absorbing ideas from other masters. He then was eligible to become a master and could apply for admission to the guild. Fees could be very high, espe-cially if an artist was not a citizen of the same city, as was the case, for example, when Petrus Christus of Baerle purchased citizenship in Bruges in order to join that city’s painters’ guild. Sometimes, an art-ist seeking admission to a guild would marry the widow of a member. (A woman could inherit her husband’s workshop, but could not run it.) Guild membership was essential for establishing an artist’s reputation and for obtaining commissions. The guild inspected paintings to evalu-ate workmanship and ensure that its members used quality materials. It also secured adequate payment for its artists’ labor. Women had far fewer opportunities than men to train as artists, in large part because of social and moral constraints that forbade women to reside as apprentices in the homes of male masters. Moreover, from the 16th century, when academic training courses supplemented and then replaced guild training, until the 20th century, women would not as a rule expect or be permitted instruction in figure painting, because it involved dissection of cadavers and study of the nude male model. Flemish women interested in pursuing art as a career—for example, Caterina van Hemessen (fig. 23-16)—most often received tutoring from fathers and husbands who were professionals and whom the women assisted in all the technical procedures of the craft. Despite these Rogier’s extraordinary ability to represent flesh and fabric rivaled Jan van Eyck’s, and few painters, not even Jan, have equaled him in rendering passionate sorrow as it vibrates through a figure or distorts a tearstained face. Rogier’s depictions of the agony of loss in Deposition and of the terror of the damned in Last Judgment (fig. 20-13a) are among the most moving and unforgettable images in religious art. It was probably Rogier whom Michelangelo had in mind when, according to the Portuguese painter Francisco de Hol-landa (1517–1584), the Italian master observed, “Flemish paint-ing [will] please the devout better than any painting of Italy, which will never cause him to shed a tear, whereas that of Flanders will cause him to shed many.”1 20-13a Van der Weyden, Last Judgment, ca. 1443–1451. 1 ft. 20-14 Petrus Christus, A Goldsmith in His Shop, 1449. Oil on wood, 39 30 × 29 100. Metropolitan Museum of Art, New York (Robert Lehman Collection, 1975). Once thought to depict Eligius, the patron saint of goldsmiths, Christus’s painting, made for the Bruges goldsmiths’ guild, is more likely a genre scene of a couple shopping for a wedding ring. obstacles, membership records of the art guilds of Bruges and other cities reveal that a substantial number of Flemish women were able to establish themselves as artists during the 15th century. That they suc-ceeded in negotiating the difficult path to acceptance as professionals is a testament to both their tenacity and their artistic skill. 580 Later Flemish painters Robert Campin, Jan van Eyck, and Rogier van der Weyden were the leading figures of the first generation of “Northern Renaissance” painters. (Art historians usually transfer to Northern Europe, with less validity than in its original usage, the term Renaissance, coined to describe the conscious revival of classical art in Italy.) The second generation of Flemish masters, active during the latter half of the 15th century, had much in common with their illustrious predeces-sors, especially a preference for using oil paints to create naturalistic representations, often, although not always, of traditional Christian subjects for installation in churches. Petrus Christus. One work of uncertain Christian content is A Goldsmith in His Shop (fig. 20-14) by Petrus Christus (ca. 1410–1472), who settled in Bruges in 1444. According to the traditional interpretation, A Goldsmith in His Shop portrays Saint Eligius (who was initially a master goldsmith before committing his life to God) sitting in his stall, showing an elegantly attired couple a selection of rings. The bride’s betrothal girdle lies on the table as a symbol of chastity, and the woman reaches for the ring the goldsmith weighs. The artist’s inclusion of a crystal container for Eucharistic wafers (on the lower shelf to the right of Saint Eligius) and the scales (a ref-erence to the Last Judgment) supports a religious interpretation of this painting and continues the Flemish tradition of imbuing every-day objects with symbolic significance. A halo once encircled the goldsmith’s head, seemingly confirming the religious nature of this scene. Scientists have determined, however, that the halo was a later addition by another artist, and restorers have removed it. Most scholars now think that the painting, although not devoid of religious content, should be seen as a vocational painting of the type often produced for installation in Flemish guild chapels. Although the couple’s presence suggests a marriage portrait, the patrons were probably not the couple portrayed but rather the goldsmiths’ guild in Bruges. Saint Eligius was the patron saint of gold-and silver-smiths, blacksmiths, and metalworkers, all of whom shared a chapel in a building adjacent to their meetinghouse. The reconsecration of this chapel took place in 1449, the same date as the Christus paint-ing. Therefore, it seems probable that the artist painted A Goldsmith in His Shop, which illustrates an economic transaction and focuses on the goldsmith’s profession, specifically for the guild chapel. A guild chapel is also the likely original setting of Rogier van der Weyden’s Saint Luke Drawing the Virgin (fig. 20-1). As noted, in the 15th century, in addition to their primary function as “trade unions,” guilds played an increasingly important role in community life and often became major patrons of art in their cities (see “The Artist’s Profession in Flanders,” page 580). Christus went to great lengths to produce a historically cred-ible image. For example, the variety of objects depicted in the paint-ing serves as advertisement for the goldsmiths’ guild. Included are the goldsmiths’ raw materials (precious stones, beads, crystal, coral, and seed pearls) scattered among finished products (rings, buckles, and brooches). The pewter vessels on the upper shelves are dona-tion pitchers, which town leaders gave to distinguished guests. All these meticulously painted objects not only attest to the centrality and importance of goldsmiths to both the secular and sacred com-munities but also enhance the naturalism of the painting. The con-vex mirror in the foreground showing another couple and a street with houses serves to extend the painting’s space into the viewer’s space, further creating the illusion of reality, as in Jan van Eyck’s Arnolfini portrait (fig. 20-11). Dieric Bouts. In Last Supper (fig. 20-15), Dieric Bouts (ca. 1415–1475) of Haarlem chose a different means of suggesting 1 ft. 20-15 Dieric Bouts, Altarpiece of the Holy Sacrament, Saint Peter’s, Louvain, Belgium, 1464–1468. Oil on wood, center panel (Last Supper) 69 × 59. The Last Supper, the central panel of this altarpiece, is one of the earliest Northern Renaissance paintings to employ Italian linear perspective. Included are portraits of the work’s Flemish patrons. spatial recession. The painting is the central panel of the Altarpiece of the Holy Sacrament, which the Confraternity of the Holy Sacra-ment in Louvain commissioned in 1464. Four years later, Bouts became the city’s offi-cial painter and was asked to produce a series of panels—including Wrongful Beheading of the Count (fig. 20-15a) and Justice of Otto III (fig. 20-15B)—for Louvain’s town hall. Bouts’s Last Supper is one of the earliest Northern Renaissance paintings to employ linear perspective and a vanishing point to create the illusion of depth. All of the central room’s orthogonals (converging diagonal lines imagined to be behind and perpendicular to the picture plane) lead to the vanishing point in the center of the mantelpiece above Jesus’s head. The small side room, however, has its own vanishing point, and neither it nor the vanishing point of the main room falls on the horizon of the landscape (the horizon line) seen through the windows, as in Italian Renaissance paintings composed according 20-15B BoutS, Justice of Otto III, ca. 1470–1473. to the strict rules of linear perspective (see “Linear Perspective,” page 599). In Last Supper, Bouts did not depict the biblical narrative itself but instead presented Jesus in the role of a priest performing the liturgical ritual of the consecration of the Eucharistic wafer. By dividing the apostles into two asymmetrical groups with no one sitting at the middle of the near side of the table, Bouts focused attention on the Savior and the food platter and wine chalice at the table’s center. Bouts’s Last Supper contrasts strongly with other depictions of the same subject, which often represented Jesus’s prediction of Judas’s betrayal or his comforting of John. The Confraternity of the Holy Sacrament dedicated itself to the worship of the Eucharist, and the smaller panels on the altarpiece’s wings depict Old Testament prefigurations of the Eucharist—for example, at the top right, the gathering of manna in the desert and, at bottom left, Jews celebrating Passover (the occasion for Jesus’s final meal). Bouts also added four servants (two looking through the window-like opening in the rear wall and two standing) not mentioned in the New Testament account, all dressed in Flemish attire. These are portraits of the four members of the confraternity who contracted Bouts to paint the altarpiece, continuing the Flemish tradition of inserting into representations of biblical events portraits of the painting’s patrons, first noted in the Mérode Triptych (fig. 20-8), where the Inghelbrechts are also looking into the room from out-side; that is, they are not participants in but witnesses of the biblical event. Hugo van der Goes. By the mid-15th century, Flemish art had achieved renown throughout Europe. The Portinari Altarpiece (fig. 20-16), for example, is a large-scale Flemish work in a family chapel in Florence, Italy. The artist who received the commission was Hugo van der Goes (ca. 1440–1482), who joined the painters’ guild of Ghent as a master in 1467 and served as the guild’s dean from 1473 to 1475. Hugo painted the triptych in Flanders for Tom-maso Portinari, an Italian shipowner and agent in Bruges for the powerful Medici bank of Florence. The altarpiece was Portinari’s gift to the church of Sant’Egidio, and he appears on the wings with his family and their patron saints, who are much larger than the donors. The main panel, Adoration of the Shepherds, depicts a sub-ject based on the 14th-century vision of a Swedish saint in which Mary, instead of cradling her newborn son, kneels solemnly to join Joseph, the angels, and the shepherds in adoring the infant Savior, who lies on the ground and glows with divine light. In order to situate the main actors at the center of the panel, Hugo tilted the ground, a compositional device he may have derived from the tilted stage floors of 15th-century mystery plays. Three shepherds enter 1 ft. 20-16 Hugo van der Goes, Adoration of the Shepherds (Portinari Altarpiece, open), from Sant’Egidio, Florence, Italy, ca. 1473–1478. Tempera and oil on wood, center panel 89 3 1 20 × 109, each wing 89 3 1 20 × 49 7 1 20. Galleria degli Uffizi, Florence. This altarpiece is a rare instance of the awarding of a major commission in Italy to a Flemish painter. The Florentines admired Hugo’s realistic details and brilliant portrayal of human character from the right rear. Hugo represented them in attitudes of wonder, piety, and gaping curiosity. Their lined faces, work-worn hands, and uncouth dress and manner indicate that they are common folk, very different than the three magi who will soon come to adore the newborn Jesus. The architecture and a continuous Northern European land-scape unify the three panels. (The bare trees of the setting estab-lish that it is winter, the season of Christ’s birth.) Symbols surface throughout the altarpiece. Iris and columbine flowers are emblems of the sorrows of the Virgin. The angels represent the 15 joys of Mary. A sheaf of wheat stands for Bethlehem (the “house of bread” in Hebrew), a reference to the Eucharist. The harp of David, embla-zoned over the building’s portal in the middle distance (just to the right of the Virgin’s head), signifies the ancestry of Christ. To stress the meaning and significance of the illustrated event, Hugo revived medieval pictorial devices. Small scenes shown in the background of the altarpiece represent (from left to right) the journey of Mary and Joseph to Bethlehem, the annunciation to the shepherds, and the arrival of the magi. Hugo’s variation in the scale of his figures to differentiate them by their importance to the central event also reflects older traditions. Still, he put a vigorous, penetrating realism to work in a new direction, characterizing human beings according to their social level while showing their common humanity. After Portinari placed the altarpiece in his family’s chapel in the Florentine church of Sant’Egidio, it created a considerable stir among Italian artists. Although the painting may have seemed unstructured to the Italians and the employing of hierarchy of scale old fashioned, Hugo’s masterful technique and what the Floren-tines deemed incredible realism in representing drapery, flowers, animals, and, above all, human character and emotion made a deep impression on them. At least one Florentine artist, Domenico Ghir-landaio (figs. 21-26 and 21-27), paid tribute to the Flemish master by using Hugo’s most striking motif, the adoring shepherds, in one of his own Nativity paintings. Hans Memling. Hugo’s contemporary, Hans Memling (ca. 1430–1494), may have trained as a painter in Rogier van der Weyden’s studio. He became a citizen of Bruges in 1465 and received numer-ous commissions from the city’s wealthy merchants, Flemish and foreign alike. He specialized in portraits of his patrons (one of whom was Tommaso Portinari; fig. 20-18A) and images of the Madonna. Memling’s many paintings of the Virgin portray young, slight, pretty princesses, each holding a doll-like infant Jesus. The center panel of the Saint John Altarpiece is Virgin with Saints and Angels (fig. 20-17). The patrons of this altarpiece—two brothers and two sisters of the order of the Hospital of Saint John in Bru-ges—appear on the exterior side panels (not illustrated). In the main panel, two angels, one playing a musical instrument and the other holding a book, flank the Virgin. To the sides of Mary’s throne stand Saints John the Baptist on the left and John the Evangelist on the right. Seated in the foreground are Saints Catherine and Barbara. This gathering celebrates the mystic marriage of Saint Catherine of Alexandria, one of many virgin saints believed to have entered into a spiritual marriage with Christ. As one of the most revered virgins of Christ, Saint Catherine provided a model of devotion that resonated with women viewers (especially nuns). The altarpiece exudes an opu-lence that results from the rich colors, the meticulously depicted tap-estries and brocades, and the serenity of the figures. The composition 1 ft. 20-17 Hans Memling, Virgin with Saints and Angels, center panel of the Saint John Altarpiece, Hospitaal Sint Jan, Bruges, Belgium, 1479. Oil on wood, center panel 59 7 3 40 × 59 7 3 40, each wing 59 7 3 40 × 29 7 1 80. Memling specialized in images of the Madonna. His Saint John Altarpiece exudes an opulence that results from the sparkling and luminous oil colors and the realistic depiction of rich tapestries and brocades 20-18 Hans Memling, Diptych of Martin van Nieuwenhove, 1487. Oil on wood, each panel 19 5 3 Bruges. 80 × 19 10. Memlingmuseum, In this diptych, the Virgin and Child pay a visit to the home of 23-year-old Martin van Nieuwenhove. A round convex mirror reflects the three figures and unites the two halves of the diptych spatially. is balanced and serene, the color sparkling and luminous, and the execution of the highest technical quality. Memling combined portraiture and Madonna imagery again in a much less ambitious—and more typical—work (fig. 20-18) that he painted for Martin van Nieuwenhove (1464–1500). The smaller scale of the private commission and its much more intimate character underscore the different approaches that painters regu-larly took when producing artworks for different kinds of patrons and for public versus private settings. Martin van Nieuwenhove was a member of an important Bruges family that held various posts in the civic government. He himself served as burgomeister of Bruges in 1497. The painting takes the form of a diptych, with the patron portrayed on the right wing praying to the Madonna and Child on the left wing, a format invented by Rogier van der Weyden. Accord-ing to inscriptions on the frames, van Nieuwenhove commissioned the work in 1487 when he was 23 years old. He died only 13 years later, 3 years after becoming mayor. The format of the van Nieuwenhove diptych follows the pat-tern that Memling used earlier for his wedding triptych Tom-maso Portinari and Maria Baroncelli (fig. 20-18a) but with only a single (male) portrait. (The lost central section of the Portinari triptych probably closely resembled the van Nieu-wenhove Madonna and Child.) Memling’s portrayals of Mary and Jesus consistently feature a tender characterization of the young Virgin and her nude infant son. Here, however, Memling set both the Madonna and her patron in the interior of a well-appointed Flemish home featuring stained-glass windows. The window to the left of the Virgin’s head bears van Nieuwenhove’s coat of arms. The window behind the donor depicts his patron saint, Martin of Tours. These precisely recorded details identify the home as van Nieuwenhove’s (the Minnewater Bridge in Bru-ges is visible through the open window), and the Madonna and Child have honored him by coming to his private residence. Even Saint Luke had to go to the Virgin’s home to paint her portrait (fig. 20-1), but the conceit is a familiar one in Flemish painting. An early example is the Annunciation taking place in the home of Peter Inghelbrecht in Robert Campin’s Mérode Triptych (fig. 20-8). In the Memling portrait diptych, the Christ Child sits on the same ledge as the donor’s open prayer book. Also uniting the two wings of the diptych is the round con-vex mirror behind the Vir-20-18a MeMling, Tommaso Porti-nari and Maria Baroncelli, ca. 1470. gin’s right shoulder in which the viewer sees the reflection of the Virgin’s back and van Nieuwenhove as well as the rest of the room (compare figs. 20-11 and 20-14). 584 FRANCE In contrast to the prosperity and peace that Flanders enjoyed dur-ing the 15th century, in France the Hundred Years’ War crippled economic enterprise and prevented political stability. The anarchy of war and the weakness of the kings gave rise to a group of duch-ies, each with significant power and the resources to commission major artworks. The strongest and wealthiest of these has already been examined—the duchy of Burgundy, which controlled Flanders, as well as the duchy of Berry in France. The French dukes of Bour-bon and Nemours as well as members of the French royal court were also important art patrons. In addition to illustrated books, such as the Très Riches Heures du Duc de Berry (figs. 20-5 and 20-6), inde-pendent paintings were popular in France, as in Flanders, for private devotional use. The preferred medium was oil paint on wood panels. Jean Fouquet. Among the French artists whose paintings were in high demand was Jean Fouquet (ca. 1425–1478). Fouquet’s workshop was in Tours, and he received important commissions from King Charles VII (r. 1422–1461, the patron and client of Jacques Coeur; fig. 13-31) and from the duke of Nemours and other elite members of the French aristocracy. Fouquet painted a diptych (fig. 20-19) for Étienne Chevalier, who, despite his lowly birth as the son of one of Charles VII’s secretaries, rose to become royal treasurer in 1452. In the left panel of the Melun Diptych (named for its original location in Melun Cathedral), Chevalier appears with his patron saint, Saint Stephen (Étienne in French). Appropriately, Fouquet’s donor portrait of Chevalier depicts his prominent patron as devout—kneeling, with hands clasped in prayer. The represen-tation of the pious donor with his standing saint recalls Flemish art, as do the three-quarter stances and the realism of Chevalier’s portrait. The artist portrayed Saint Stephen, whose head also has a portraitlike quality, holding the stone symbolizing his martyrdom (death by stoning) atop a volume of the holy scriptures, thereby ensuring that viewers can properly identify the saint. Fouquet ren-dered the entire image in meticulous detail and included a highly ornamented architectural setting. In its original diptych form (the two panels are now in differ-ent museums), the viewer would follow the gaze of Chevalier and Saint Stephen over to the right panel, which depicts the Virgin Mary and Christ Child in a most unusual way—with marblelike flesh, sur-rounded by red and blue cherubs (chubby winged child angels). The juxtaposition of these two images enabled the patron to bear wit-ness to the sacred. The integration of sacred and secular (especially the political or personal), prevalent in other Northern Renaissance artworks, also emerges here, which complicates the reading of this diptych. Agnès Sorel (1421–1450), the mistress of Charles VII, was Fouquet’s model for the Virgin Mary, whose left breast is exposed, presumably to nurse the infant Jesus, but she does not look at her son or the viewer. (Mary as nursing mother is a not uncommon way of representing the Virgin in Northern Europe; compare fig. 20-1.) Chevalier commissioned this painting after Sorel’s death, probably by poisoning while pregnant with the king’s child. Thus, in addition to the religious interpretation of this diptych, there is surely a personal 1 ft. 1 ft. 20-19 Jean Fouquet, Melun Diptych, ca. 1452. Left wing: Étienne Chevalier and Saint Stephen. Oil on wood, 39 1 Gemäldegalerie, Staatliche Museen zu Berlin, Berlin. Right wing: Virgin and Child. Oil on wood, 39 11 Museum voor Schone Kunsten, Antwerp. 40 × 29 9 1 20 × 29 9 1 20. 20. Koninklijk Fouquet’s meticulous representation of a pious kneeling donor with a standing patron saint recalls Flemish painting, as do the three-quarter stances and the realism of the donor’s portrait head 20-20 Konrad Witz, Miraculous Draft of Fish, exterior wing of Altarpiece of Saint Peter, from the Chapel of Notre-Dame des Maccabées, Cathedral of Saint Peter, Geneva, Switzerland, 1444. Oil on wood, 49 30 × 59 10. Musée d’Art et d’Histoire, Geneva. Konrad Witz set this biblical story on Lake Geneva. The painting is one of the first 15th-century works depicting a specific locale and is noteworthy for the painter’s skill in rendering water effects. 1 ft. and political narrative here as well, made all the more complex given that Sorel gave birth to three of the king’s daughters, but never a son. HOLY ROMAN EMPIRE Because the Holy Roman Empire (whose core was Germany) did not participate in the drawn-out saga of the Hundred Years’ War, its economy remained stable and prosperous. Without a domi-nant court to commission artworks, wealthy merchants and clergy became the primary German art patrons during the 15th century. panel painting The art of the early Northern Renaissance in the Holy Roman Empire displays a pronounced stylistic diversity. Some artists fol-lowed developments in Flemish painting, and large-scale altar-pieces featuring naturalistically painted biblical themes were familiar sights in German churches. Konrad Witz. Among the most notable 15th-century German altarpieces is the Altarpiece of Saint Peter, painted by Konrad Witz (ca. 1400–1446) for the chapel of Notre-Dame des Maccabées in the Cathedral of Saint Peter in Geneva. Signed by “Magister Con-radus of Basel” and dated 1444, the altarpiece is a triptych featuring Witz’s paintings on both the exterior and interior. Miraculous Draft of Fish (fig. 20-20) appears on one of the exterior wings. The other (not illustrated) depicts the release of Saint Peter from prison. The central panel is lost. On the interior wings, Witz painted scenes of the adoration of the magi and of Saint Peter’s presentation of the donor (Bishop François de Mies) to the Virgin and Child. Miraculous Draft of Fish shows Peter, the first pope, unsuccessfully trying to emulate Christ walking on water. The choice of this episode from among oth-ers in Peter’s life (see “Early Christian Saints,” pages 246–247) may be a commentary on the part of Witz’s patron, the Geneva cardinal, on the limited power of the pope in Rome. The painting is particularly significant because of the land-scape’s prominence. Witz showed his considerable skill in the recording of water effects—the shimmering sky on the slowly mov-ing lake surface, the mirrored reflections of the figures in the boat, and the transparency of the shallow water in the foreground. He observed and represented the landscape so carefully that art histo-rians have been able to determine the exact location shown. Witz presented a view of the shores of Lake Geneva, with the town of Geneva on the right and Le Môle Mountain in the dis-tance behind Christ’s head. This painting is one of the first 15th-century works depict-ing a specific, identifiable site. The work of other leading German painters of the mid-15th century—for exam-ple, Stefan Lochner (ca. 1400–1451)—retained medieval features to a much greater degree, as is immediately evident in a comparison between Witz’s landscape and Lochner’s Madonna in the Rose Gar-den (fig. 20-20a). 586 CHAPTER 20 Late Medieval and Early Renaissance Northern Europe Copyright 2021 Cengage Learning. All Rights Reserved. May not be copied, scanned, or duplicated, in whole or in part. Due to electronic rights, some third party content may be suppressed from the eBook and/or eChapter(s). Editorial review has deemed that any suppressed content does not materially affect the overall learning experience. Cengage Learning reserves the right to remove additional content at any time if subsequent rights restrictions require it. 20-20a lochner, Madonna in the Rose Garden, ca. 1435–1440. sculpture As in Flanders (fig. 20-3), in the Holy Roman Empire many of the leading 15th-century artists specialized in carving large wood retables. These grandiose sculpted altarpieces reveal the lingering power of the Late Gothic style. Veit Stoss. The sculptor Veit Stoss (ca. 1447–1533) trained in the Upper Rhine region, possibly in Strasbourg, but settled in Kraków (in present-day Poland) in 1477. In that year, he began work on a monumental altarpiece for the church of Saint Mary in Kraków, paid for by the city’s citizens. In the central boxlike shrine is Death and Assumption of the Virgin (fig. 20-21), peopled by huge carved, painted, and gilded figures, some 9 feet high. On both the outer and inner faces of the wings, Stoss portrayed scenes from the lives of Christ and Mary. The altar forcefully expresses the intense piety of Gothic culture in its late phase, when artists used every figural and ornamental motif in the repertoire of Gothic art to heighten the emotion and glorify sacred events. In the center panel of the Kraków altarpiece, Christ’s disciples congregate around the Virgin, who collapses, dying. One of them supports her, while another wrings his hands in grief. Above, Mary rises to Heaven with the resurrected Christ. Stoss posed others in attitudes of woe and psychic shock, striving for realism in every minute detail. He engulfed the figures in restless, twisting, and curving swaths of drapery whose broken and writhing lines unite the whole tableau in a vision of agitated emotion. The artist’s massing of sharp, broken, and pierced forms that dart flamelike through the composition—at once unifying and animating it—recalls the design principles of Late Gothic architecture (fig. 13-28). Indeed, in the Kraków altar-piece, Stoss merged sculpture and architecture, enhancing their union with paint and gilding. The Kraków altarpiece gives modern viewers the opportunity to appreciate the polychromy that was standard in ancient and medieval sculpture in both wood and stone. Examples such as this one are unfortunately rare. The vast majority of preserved statues and reliefs from antiquity and the Middle Ages give a false impres-sion of their original appearance (compare figs. 3-13A and 5-63A) because the paint has disappeared. 10 ft. 20-21 Veit Stoss, Death and Assumption of the Virgin, center panel of the Altarpiece of the Virgin Mary (with wings open), Church of Saint Mary, Kraków, Poland, 1477–1489. Painted and gilded lime-wood, central panel 239 90 high. In this huge painted and gilded sculptured wood altarpiece, Stoss used every figural and ornamental element from the vocabulary of Gothic art to heighten the emotion and glorify the sacred event. 20-22 Tilman Riemenschneider, Assumption of the Virgin (Creglingen Altarpiece), Herrgottskirche, Creglingen, Germany, ca. 1495–1505. Lime-wood, 309 20 high. Riemenschneider specialized in carving large unpainted wood retables. His works feature intricate Gothic trac-ery and religious figures whose bodies are almost lost within their swirling garments. into fluid motion by employing an endless and restless line running through the garments of the figures. The draperies float and flow around bodies lost within them, serving not as descrip-tions but as design elements that tie the figures to one another and to the framework. A look of psychic strain, a facial expression common in Riemenschneider’s work, heightens the spiritu-ality of the figures, immaterial and weightless as they appear. Graphic arts 10 ft. A new age blossomed in the 15th century with a sudden technological advance that had wide-spread effects—the invention by Johannes Gutenberg (ca. 1400–1468) of moveable type around 1450 and the development of the print-ing press. Printing had been known in China centuries before, but had never fostered, as it did in 15th-century Europe, a revolution in written communication and in the generation and man-agement of information. Printing provided new and challenging media for artists, and the earli-est form was the woodcut. Artists produced inex-pensive woodcuts such as the Buxheim Saint Christopher (fig. 20-22a) before the development of moveable-type print-ing. But when the im-proved economy and a rise in literacy necessi-tated the production of illustrated books on a grand scale, artists met the challenge of bringing the woodcut picture onto the same letterpress text pages (see “How to Illustrate Printed Books,” page 589). 20-22a Buxheim Saint Christopher, 1423(?). Martin Schongauer. The woodcut medium hardly had matured when printmakers intro-duced the technique of engraving (see “Engrav-Tilman Riemenschneider. Assumption of the Virgin is also the center panel of the Creglingen Altarpiece (fig. 20-22), carved by Tilman Riemenschneider (ca. 1460–1531) of Würzburg for a parish church in Creglingen, Germany. Riemenschneider incor-porated intricate Gothic forms, especially in the 30-foot-tall altar-piece’s elaborate canopy, but unlike Stoss, he did not paint the figures or the background, relying instead purely on sculptural form. The Würzburg master succeeded in setting the whole design ing and Etching,” page 590). Begun in the 1430s and well developed by 1450, engraving proved much more flexible than woodcut. Predictably, in the second half of the century, engraving began to replace the woodcut process for making both book illustrations and widely popular single prints. Not surprisingly, many of the earli-est engravers were professional goldsmiths, who easily applied their training in incising designs into gold and silver jewelry to the new art form. how to illustrate printed Books With the invention of moveable type in the 15th century and the new widespread availability of paper from commercial mills, the art of print-making developed rapidly in Europe. A print is an artwork created by transferring a design on a metal plate or a wood or stone block to a sheet of paper. Prints are usually produced in multiple impressions. The set of prints that an artist creates from a single block or plate is called an edition. Like the manufacturing of books on a press, the printmaking process involves the transfer of ink from a printing surface to paper. This can be accomplished in several ways. The oldest and simplest is the relief method—carving the design into a surface, usually a soft wood, such as pear. Relief print-ing requires artists to conceptualize their images in reverse—that is, they must draw mirror images of the intended design on the wood block. They then remove the surface areas around the drawn lines using a gouging instrument. Thus when the printmaker inks the ridges that correspond to the original (reversed) drawing, the hollow areas remain dry, and a correctly oriented image results when the artist presses the printing block against paper. Because artists produce woodcuts through a subtractive process (removing parts of the mate-rial), it is difficult to create very thin, fluid, and closely spaced lines. Moreover, if the ridges are too thin, they will break during printing. As a result, woodcut prints (for example, figS. I-9, 20-22A, and 20-23) tend to exhibit stark contrasts between the dark lines and the blank background. During the later 15th century, publishers realized that the wood-cut was the ideal medium for solving the problem of illustrating their new printed books. The new technology quickly led to the demise of the manuscript illuminator as well as the scribe, both of whom had been essential for the production of illustrated books through the late Middle Ages, whether they were monks working in monastic scriptoria or lay professionals employed in the for-profit workshops of Paris and other cities. The most ambitious early effort to produce low-cost mass-produced printed books was the so-called Nuremberg Chronicle, a history of the world produced in Nuremberg by anton koBerger (ca. 1445–1513), with more than 650 illustrations furnished by the workshop of Michael WolgeMut (1434–1519). The hand-colored page illustrated here (fig. 20-23) represents Madeburga (modern Magde-burg, Germany). The blunt, simple lines of the woodcut technique give a detailed perspective of Magdeburg, its harbor and shipping, its walls and towers, its churches and municipal buildings, and a statue-topped columnar monument. Despite the numerous architectural structures, this illustration is not an accurate depiction of the city but the prod-uct of the artist’s fanciful imagination. Wolgemut often used the same image to illustrate different cities, even those with very little in common—for example, Verona, Italy, and Damascus, Syria. This depiction of Magdeburg is a generic view of a late medieval German town, not a portrait of a specific place. Regardless, the Nuremberg Chronicle is a monument to a new craft, which expanded in concert with the art of the printed book. 20-23 Michael Wolgemut and workshop, Madeburga, page from the Nurem-berg Chronicle, 1493. Woodcut, 19 6 1 80 × 29 15 80. Printed by Anton Koberger. The Nuremberg Chronicle is an early example of wood-cut illustrations in printed books. The more than 650 pictures include detailed views of towns, but they are generic rather than specific portrayals. 1 engraving and etching In contrast to the production of relief prints, in the intaglio method of printmaking, the artist incises (cuts) the lines on a metal plate, often copper, rather than cuts away the area around the lines, as the relief printer does. The image can be created on the plate manually (engrav-ing or drypoint; for example, fig. 20-24) using a tool (a burin or stylus) or chemically (etching; for example, fig. 25-16). In the etching process, an acid bath eats into the exposed parts of the plate where the artist has drawn through an acid-resistant coating. When the artist inks the surface of the intaglio plate and wipes it clean, the ink is forced into the incisions. Then the artist runs the plate and paper through a roller press, and the paper absorbs the remaining ink, creating the print. Because the artist “draws” the image onto the plate, intaglio prints differ in char-acter from relief prints, such as woodcuts. Engravings, drypoints, and etchings generally present a wider variety of linear effects, as is imme-diately evident in a comparison of the roughly contemporaneous wood-cut (fig. 20-23) by Michael Wolgemut and an engraving (fig. 20-24) by Martin Schongauer. Intaglio prints also often reveal to a greater extent evidence of the artist’s touch, the result of the hand’s changing pressure and shifting directions. The paper and inks that artists use also affect the finished look of the printed image. During the 15th and 16th centuries, European print-makers used papers produced from cotton and linen rags that paper-makers mashed with water into a pulp. The papermakers then applied a thin layer of this pulp to a wire screen and let it dry to create the paper. As contact with Asia increased, printmakers made greater use of what was called Japan paper (of mulberry fibers) and China paper. Artists, then as now, could select from a wide variety of inks. The type and pro-portion of the ink ingredients affect the consistency, color, and oiliness of inks, which various papers absorb differently. Paper is lightweight, and the portability of prints has been an important factor in their appeal to artists. The opportunity to produce numerous impressions from the same print surface also made printmak-ing attractive to 15th-and 16th-century artists. In addition, prints can be sold at much lower prices than paintings or sculptures. Consequently, prints reached a far wider audience than did one-of-a-kind artworks. The number and quality of existing 15th-and 16th-century European prints attest to the importance of the new print medium. 20-24 Martin Schongauer, Saint Anthony Tormented by Demons, ca. 1480. Engraving, 19 1 40 × 90. Fondazione Magnani Rocca, Corte di Mamiano. Schongauer was the most skilled of the early masters of metal engraving. By using a burin to incise lines in a copper plate, he was able to create a marvelous variety of tonal values and textures. 1 in. Martin Schongauer (ca. 1430–1491) of Colmar, a painter and the son of a goldsmith, was the most skilled and subtle North-ern European master of metal engraving. His Saint Anthony Tor-mented by Demons (fig. 20-24) shows both the versatility of the medium and the artist’s mastery of it. The stoic saint is caught in a revolving thornbush of spiky demons, who claw and tear at him furiously. With unsurpassed skill and subtlety, Schongauer incised lines of varying thickness and density into a metal plate and cre-ated marvelous distinctions of tonal values and textures—from smooth skin to rough cloth, from the furry and feathery to the hairy and scaly. The use of cross-hatching (sets of engraved lines at right angles) to describe forms, which Schongauer probably devel-oped, became standard among German graphic artists. The Italians preferred parallel hatching (fig. 21-29) and rarely adopted cross-hatching, which, in keeping with the general Northern Renaissance approach to art, tends to describe the surfaces of things rather than their underlying structures. Schongauer probably engraved Saint Anthony around 1480. By then, the political geography of Europe had changed dramatically. Charles the Bold, who had assumed the title of duke of Burgundy in 1467, died in 1477, bringing to an end the Burgundian dream of forming a strong middle kingdom between France and the Holy Roman Empire. After Charles’s death at the battle of Nancy, the French monarchy reabsorbed the southern Burgundian lands, and the Netherlands passed to the Holy Roman Empire by virtue of the dynastic marriage of Charles’s daughter, Mary of Burgundy (fig. 20-7), to Maximilian of Habsburg, inaugurating a new political and artistic era in Northern Europe, examined in Chapter 23. The next two chapters, however, explore Italian developments in painting, sculpture, and architecture during the 15th and 16th centuries. 590 French ducal courts ■ The most powerful rulers north of the Alps during the first three quarters of the 15th century were the dukes of Burgundy. They controlled Flanders, which derived its wealth from wool and banking, and were great art patrons. Duke Philip the Bold (r. 1363–1404) endowed the Carthusian monastery at Champmol, near Dijon, which became a ducal mausoleum. The head of Philip’s sculptural workshop was Claus Sluter, whose Well of Moses features innovative statues of prophets with portraitlike features and realistic costumes. ■ In the 15th century, Northern European artists popularized the use of oil paints on wood panels. By superimposing translucent glazes, painters could create richer colors than possible using tempera or fresco. One of the earliest examples of oil painting is Melchior Broederlam’s wings of the Retable de Champmol, painted for Philip the Bold. Carved wood narrative reliefs by Jacques de Baerze fill the retable’s interior. ■ The Limbourg brothers expanded the illusionistic capabilities of manuscript illumination in the Book of Hours they produced for Jean, duke of Berry (r. 1360–1416) and brother of King Charles V (r. 1364–1380). Their full-page calendar pictures alternately represent the nobility and the peasantry, always in seasonal, naturalistic settings with realistically painted figures. Flanders ■ A major art form in Flemish churches and private homes alike was the altarpiece with folding wings. In Robert Campin’s Mérode Triptych, the Annunciation takes place in a Flemish home. As is typical of “Northern Renaissance” art, the everyday objects in the triptych have symbolic significance. The largest altarpiece of the era is the Ghent Altarpiece, begun by Hubert van Eyck and completed by his brother Jan. ■ Jan van Eyck, Rogier van der Weyden, and others established portraiture as an important art form in 15th-century Flanders. Their subjects were successful businessmen, both Flemish and foreign, and occasionally themselves. Jan portrayed himself wearing a red turban, and Rogier represented himself in the guise of Saint Luke drawing the Virgin. ■ Among the other major Flemish painters were Petrus Christus and Hans Memling of Bruges, Dieric Bouts of Louvain, and Hugo van der Goes of Ghent, all of whom produced both altarpieces for churches and portraits for the homes of wealthy merchants. van Eyck, Ghent Altarpiece, ca. 1423–1432 de Baerze, Retable de Champmol, 1399 Limbourg brothers, Les Très Riches Heures du Duc de Berry, 1413–1416 France and the holy roman empire ■ During the 15th century, the Hundred Years’ War crippled the French economy, but dukes and mem-bers of the royal court still commissioned some notable artworks. French court art—for example, Jean Fouquet’s Melun Diptych—owes a large debt to Flemish painting in style and technique as well as in the integration of sacred and secular themes. ■ The Late Gothic style remained popular in 15th-century Germany for large carved wood retables fea-turing highly emotive figures amid Gothic tracery. The leading sculptors were Veit Stoss and Tilman Riemenschneider. ■ The major German innovation of the 15th century was the development of the printing press, which publishers soon used to produce books with woodcut illustrations. Woodcuts are relief prints in which the artist carves out the areas around the lines to be printed. ■ German artists, such as Martin Schongauer, were the earliest masters of engraving. The intaglio technique enables a wider variety of linear effects because the artist incises the image directly onto a metal plate. Nuremberg Chronicle, 1493 Fouquet 21-1a Inspired by a poem by Angelo Poliziano, Botticelli painted Birth of Venus for the Medici between 1484 and 1486. At the left, Zephyrus, carrying Chloris, blows Venus on a cockleshell to Cyprus.  21-1b Botticelli’s revival of the theme of the female nude, largely absent from medieval art, was consistent with the Quattrocento view that beholding physical beauty prompts the contemplation of spiritual beauty. 1 ft. 21-1 Sandro Botticelli, Birth of Venus, ca. 1484–1486. Tempera on canvas, 59 90 × 99 20. Galleria degli Uffizi, Florence.  21-1c Awaiting the newborn goddess of love on her sacred island is the nymph Pomona, who runs to meet Venus with a brocaded mantle. Her draperies undulate loosely in the gentle gusts of wind. Copyright

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Analysis of 'The Little Seed'

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8.4 and 8.5 Enzymes speeding up processes and regulation of enzymes

Seed Germination, Dispersal, and Plant Cultivation

BIOL 101 Complete Study Guide Unit 1: DNA, RNA, and Genetics 1. DNA structure What is DNA? DNA stands for deoxyribonucleic acid. It stores the genetic information that tells cells how to function, grow, and reproduce. What is the shape of DNA? DNA has a double-helix shape, which looks like a twisted ladder. What are the building blocks of DNA? Nucleotides. What are the three parts of a DNA nucleotide? * A phosphate group * A five-carbon sugar called deoxyribose * A nitrogenous base What are the four nitrogenous bases in DNA? * Adenine (A) * Thymine (T) * Cytosine (C) * Guanine (G) What are the DNA base-pairing rules? * Adenine pairs with Thymine (A–T). * Cytosine pairs with Guanine (C–G). What does complementary base pairing mean? Each DNA base has a specific partner. Knowing one DNA strand allows you to determine the matching strand. Example: * Original: A T C G * Complement: T A G C What holds the two DNA strands together? Hydrogen bonds between the nitrogenous bases. 2. DNA vs. RNA What is RNA? RNA stands for ribonucleic acid. It helps use the genetic instructions stored in DNA to make proteins. What are the main differences between DNA and RNA? DNA RNA Usually double-stranded Usually single-stranded Contains deoxyribose sugar Contains ribose sugar Uses thymine (T) Uses uracil (U) Stores genetic information Helps make proteins What are the four bases in RNA? * Adenine (A) * Uracil (U) * Cytosine (C) * Guanine (G) What replaces thymine in RNA? Uracil. What are the three main types of RNA? * mRNA (messenger RNA): Carries genetic instructions from DNA to the ribosome. * tRNA (transfer RNA): Brings amino acids to the ribosome. * rRNA (ribosomal RNA): Makes up part of the ribosome and helps build proteins. 3. DNA replication What is DNA replication? DNA replication is the process of making an identical copy of DNA before a cell divides. Why is DNA replication important? It ensures that each new cell receives a copy of the genetic information. What does semiconservative replication mean? Each new DNA molecule contains one original strand and one newly made strand. What enzyme unzips DNA? Helicase. What enzyme adds new DNA nucleotides? DNA polymerase. What enzyme connects DNA fragments on the lagging strand? DNA ligase. What is the leading strand? The strand that is copied continuously in the direction of the replication fork. What is the lagging strand? The strand that is copied in short sections called Okazaki fragments. What are Okazaki fragments? Short pieces of newly made DNA that form on the lagging strand. What is the job of DNA ligase? It joins the Okazaki fragments together. What are telomeres? Protective sections of DNA found at the ends of chromosomes. What is the purpose of telomeres? They help protect important genetic information from being lost when DNA is copied. What is DNA repair? The process cells use to identify and correct mistakes or damage in DNA. 4. Transcription What is transcription? Transcription is the process of copying genetic information from DNA into mRNA. Where does transcription happen in eukaryotic cells? In the nucleus. What enzyme helps make RNA during transcription? RNA polymerase. What is the purpose of transcription? To create an mRNA copy of a gene that can be used to make a protein. What are the RNA base-pairing rules? * DNA A pairs with RNA U. * DNA T pairs with RNA A. * DNA C pairs with RNA G. * DNA G pairs with RNA C. Example: DNA: A T C G RNA: U A G C What happens to mRNA before it leaves the nucleus in eukaryotic cells? It undergoes RNA processing. What are introns? Sections of RNA that are removed during RNA processing. What are exons? Sections that remain and are joined together to form mature mRNA. What is RNA splicing? The removal of introns and joining of exons. 5. Translation What is translation? Translation is the process of using mRNA instructions to build a protein. Where does translation happen? At the ribosomes in the cytoplasm or on the rough endoplasmic reticulum. What is a codon? A group of three mRNA bases that codes for an amino acid or a stop signal. What is an anticodon? A group of three bases on tRNA that pairs with a matching mRNA codon. What is the role of tRNA? It carries amino acids to the ribosome. What is the role of the ribosome? It reads mRNA and connects amino acids to build a protein. What is the start codon? AUG. What amino acid does AUG code for? Methionine. What are the three stop codons? * UAA * UAG * UGA What happens when a stop codon is reached? Translation ends, and the completed protein is released. 6. Gene expression and regulation What is gene expression? Gene expression is the process of using information from a gene to make a functional product, such as a protein. What is gene regulation? Gene regulation controls when, where, and how much a gene is expressed. Why is gene regulation important? It allows cells to perform different functions even though many cells contain the same DNA. What is an operon? A group of genes controlled together, commonly found in bacteria. What is the lac operon? A group of bacterial genes involved in breaking down lactose. What happens when lactose is present? Lactose helps remove the repressor’s blockage, allowing the genes to be expressed. What happens when lactose is absent? The repressor binds to the operator and prevents the genes from being expressed. Unit 2: Evolution and Natural Selection 1. Understanding evolution What is evolution? Evolution is the change in inherited characteristics of a population over generations. Does an individual organism evolve during its lifetime? No. Populations evolve over generations. What is a population? A group of individuals of the same species living in the same area. What is a species? A group of organisms that can reproduce with one another and produce fertile offspring, under the biological species concept. What is natural selection? Natural selection is the process in which organisms with inherited traits that improve survival or reproduction tend to leave more offspring. Who is associated with the theory of evolution by natural selection? Charles Darwin. What is adaptation? An inherited characteristic that helps an organism survive or reproduce in its environment. What is biological fitness? An organism’s ability to survive and reproduce, measured by its contribution of offspring to the next generation. Does the strongest organism always have the highest fitness? No. Fitness depends on how well an organism’s inherited traits help it reproduce in its particular environment. 2. Natural selection What are the main requirements for natural selection? * Individuals in a population have variation. * Some variation is inherited. * Organisms produce more offspring than can survive. * Individuals with certain inherited traits reproduce more successfully. What happens to helpful inherited traits over generations? They may become more common in the population. Can natural selection create traits because organisms need them? No. Natural selection acts on existing inherited variation. It does not create a needed trait simply because an organism needs it. What is an example of natural selection? If insects vary in color and birds more easily spot one color, insects with the less visible color may survive and reproduce more often. Over generations, that color may become more common. 3. Population genetics What is an allele? An alternative version of a gene. What is a gene pool? All the alleles present in a population. What is allele frequency? The proportion of a particular allele in a population. What does it mean when allele frequencies change? The population is evolving. What is genetic variation? Differences in genetic information among individuals in a population. Why is genetic variation important? It provides the differences that natural selection can act on. 4. Mechanisms of evolution What are the five major mechanisms that can change allele frequencies? * Mutation * Gene flow * Genetic drift * Natural selection * Nonrandom mating can change genotype frequencies and, in some circumstances, influence allele frequencies indirectly through selection. What is mutation? A change in DNA sequence. How can mutations affect evolution? They can introduce new genetic variations and sometimes create new alleles. What is gene flow? The movement of alleles between populations when individuals or their reproductive cells move between them. What is genetic drift? A random change in allele frequencies, especially noticeable in small populations. What is the bottleneck effect? A type of genetic drift that happens when a population becomes much smaller due to a random event. What is the founder effect? A type of genetic drift that occurs when a small group separates from a larger population and starts a new population. What is the difference between natural selection and genetic drift? Natural selection is based on differences in reproductive success associated with inherited traits. Genetic drift changes allele frequencies randomly. 5. Hardy-Weinberg equilibrium What is Hardy-Weinberg equilibrium? A model that describes a population that is not evolving with respect to the gene being studied. What are the five Hardy-Weinberg conditions? * No mutations * Random mating * No natural selection * Very large population * No gene flow What is the Hardy-Weinberg equation? p+q=1 What does p represent? The frequency of one allele. What does q represent? The frequency of the other allele. What is the genotype frequency equation? p^2+2pq+q^2=1 What does each part represent? * : Homozygous dominant * : Heterozygous * : Homozygous recessive What does homozygous mean? Having two identical alleles for a gene. What does heterozygous mean? Having two different alleles for a gene. Practice question: If q = 0.4, what is p? Answer: p=1-0.4=0.6 Practice question: If p = 0.6 and q = 0.4, what is the frequency of homozygous dominant individuals? Answer: p^2=0.6^2=0.36 Practice question: If p = 0.6 and q = 0.4, what is the frequency of heterozygous individuals? Answer: 2pq=2(0.6)(0.4)=0.48 Practice question: If p = 0.6 and q = 0.4, what is the frequency of homozygous recessive individuals? Answer: q^2=0.4^2=0.16 6. Evidence for evolution What are some forms of evidence for evolution? * Fossil records * Comparative anatomy * Molecular similarities * Embryology * Biogeography What are fossils? Preserved remains or traces of organisms that lived in the past. What is homologous anatomy? Similar body structures inherited from a common ancestor, even if they serve different functions. What is an example of homologous structures? The forelimbs of humans, whales, bats, and cats. What is analogous anatomy? Structures that perform similar functions but evolved independently and do not come from the same ancestral structure. What is an example of analogous structures? The wings of birds and insects. What is biogeography? The study of where organisms live and how their geographic distribution relates to evolution. How can DNA provide evidence for evolution? Organisms with more similar DNA sequences often share a more recent common ancestor. 7. Speciation What is speciation? The formation of new species. What is reproductive isolation? When populations can no longer successfully reproduce with one another. What is geographic isolation? When physical barriers separate populations, such as mountains, rivers, or oceans. What is allopatric speciation? Speciation that occurs when populations are geographically separated. What is sympatric speciation? Speciation that occurs without geographic separation. Unit 3: Classification, Taxonomy, and Phylogeny 1. Classification What is taxonomy? The science of naming and classifying organisms. What is systematics? The study of biological diversity and evolutionary relationships. What is binomial nomenclature? The two-part scientific naming system for organisms. What are the two parts of a scientific name? Genus and species. How should scientific names be written? The genus is capitalized, the species name is lowercase, and both are italicized. Example: Homo sapiens. What are the main taxonomic levels from broadest to most specific? * Domain * Kingdom * Phylum * Class * Order * Family * Genus * Species What are the three domains of life? * Bacteria * Archaea * Eukarya 2. Phylogenetic trees What is a phylogenetic tree? A diagram showing evolutionary relationships among organisms. What does a branch point represent? A common ancestor where evolutionary lineages split. What is a common ancestor? An ancestral organism or population from which two or more groups evolved. What are sister taxa? Two groups that share an immediate common ancestor. What does a node represent on a phylogenetic tree? A branching point representing a common ancestor. Does being next to another organism on a tree always mean they are closely related? No. Relationships are determined by shared branch points, not simply by how close the names appear. What is a clade? An ancestor and all of its descendants. What is a derived trait? A trait that evolved in a particular lineage and can help identify relationships among organisms. Unit 4: Prokaryotes, Eukaryotes, Bacteria, and Archaea 1. Cell types What are prokaryotic cells? Cells that do not have a nucleus or membrane-bound organelles. What are eukaryotic cells? Cells that have a nucleus and membrane-bound organelles. What organisms are prokaryotes? Bacteria and Archaea. What organisms are eukaryotes? Animals, plants, fungi, and protists. What is the main difference between prokaryotic and eukaryotic cells? Eukaryotic cells have a nucleus and membrane-bound organelles. Prokaryotic cells do not. Where is DNA located in a prokaryotic cell? In a region called the nucleoid. What is the function of the cell membrane? It controls what enters and leaves the cell. What is the function of ribosomes? They make proteins. 2. Bacteria and Archaea What are bacteria? Single-celled prokaryotic organisms found in many environments. What are Archaea? Single-celled prokaryotic organisms that are genetically and chemically distinct from bacteria. What is binary fission? A form of asexual reproduction in which one cell divides into two cells. Do bacteria and archaea have a nucleus? No. What is a bacterial cell wall commonly made of? Peptidoglycan. What is peptidoglycan? A material that provides structure and support to bacterial cell walls. 3. Gram-positive and Gram-negative bacteria What are Gram-positive bacteria? Bacteria with a thick peptidoglycan cell wall that usually stain purple during Gram staining. What are Gram-negative bacteria? Bacteria with a thinner peptidoglycan layer and an outer membrane that usually stain pink or red during Gram staining. What is the major difference between Gram-positive and Gram-negative bacteria? Gram-positive bacteria have thick peptidoglycan walls, while Gram-negative bacteria have thin peptidoglycan walls and an outer membrane. 4. Endosymbiotic theory What is the endosymbiotic theory? The theory that mitochondria and chloroplasts originated from free-living bacteria that were taken into ancestral eukaryotic cells. What organelles are explained by endosymbiotic theory? Mitochondria and chloroplasts. What evidence supports endosymbiotic theory? * They contain their own DNA. * They have their own ribosomes. * They divide in a way similar to bacteria. * They have double membranes. What is the function of mitochondria? They produce most of the cell’s ATP through cellular respiration. What is the function of chloroplasts? They carry out photosynthesis in plants and some other organisms. 5. Protists and fungi What are protists? A diverse group of eukaryotic organisms that do not fit into the traditional plant, animal, or fungi kingdoms. Are all protists single-celled? No. Many are single-celled, but some are multicellular. What are fungi? Eukaryotic organisms that absorb nutrients from their surroundings. What is the cell wall of fungi made of? Chitin. What are examples of fungi? * Mushrooms * Molds * Yeasts How do fungi obtain nutrients? They release digestive enzymes into their surroundings and absorb the broken-down nutrients. What is the role of decomposers? They break down dead organisms and recycle nutrients into the environment. Unit 5: Plants 1. Plant characteristics What are the major characteristics of plants? Plants are multicellular eukaryotes that generally have chloroplasts, cellulose cell walls, and carry out photosynthesis. What is photosynthesis? The process of using light energy to convert carbon dioxide and water into sugars and oxygen. What is the function of chlorophyll? It absorbs light energy for photosynthesis. What is the function of roots? They absorb water and minerals and help anchor the plant. What is the function of stems? They support the plant and transport materials. What is the function of leaves? They are the main organs where photosynthesis occurs in most plants. 2. Alternation of generations What is alternation of generations? A plant life cycle that alternates between two multicellular stages: the haploid gametophyte and the diploid sporophyte. What is a gametophyte? The haploid stage that produces gametes. What is a sporophyte? The diploid stage that produces spores through meiosis. What does haploid mean? Having one set of chromosomes, represented by . What does diploid mean? Having two sets of chromosomes, represented by . What is fertilization? The joining of two gametes to form a diploid zygote. What is meiosis? A type of cell division that reduces the chromosome number by half and produces haploid cells. What is the difference between spores and gametes? Spores can develop into a new organism without fusing with another cell. Gametes must fuse during fertilization to form a zygote. 3. Major plant groups What are the major groups of plants? * Nonvascular plants * Seedless vascular plants * Gymnosperms * Angiosperms What are nonvascular plants? Plants that lack specialized vascular tissues for transporting water and nutrients. What are examples of nonvascular plants? Mosses and liverworts. What are vascular plants? Plants that have specialized tissues to transport water, minerals, and sugars. What are the two major vascular tissues? * Xylem * Phloem What is the function of xylem? It transports water and minerals from the roots toward the rest of the plant. What is the function of phloem? It transports sugars and other organic nutrients throughout the plant. What are seedless vascular plants? Vascular plants that reproduce using spores instead of seeds. What is an example of a seedless vascular plant? Ferns. What are gymnosperms? Seed-producing plants with seeds that are not enclosed within fruits. What is an example of a gymnosperm? Pine trees. What are angiosperms? Flowering plants that produce seeds enclosed within fruits. What is the main difference between gymnosperms and angiosperms? Gymnosperms have exposed seeds, while angiosperms produce seeds enclosed in fruits. 4. Plant reproduction What is pollination? The transfer of pollen from the male reproductive structure to the female reproductive structure of a plant. What is pollen? A structure that carries the male gametophyte in seed plants. What is the purpose of flowers? Flowers are reproductive structures of angiosperms. What develops into a fruit? The ovary of a flower. What develops into a seed? The ovule after fertilization. What is the purpose of seeds? They protect and provide resources for the developing plant embryo. Unit 6: Animals 1. Animal characteristics What are the major characteristics of animals? Animals are multicellular eukaryotes that obtain nutrients by consuming other organisms or organic material. They do not have cell walls. Are animals prokaryotic or eukaryotic? Eukaryotic. Are animals unicellular or multicellular? Multicellular. Are animals autotrophs or heterotrophs? Heterotrophs. What is a heterotroph? An organism that obtains energy and nutrients by consuming other organisms or organic material. 2. Animal body plans What is symmetry? The arrangement of body parts around an axis or center. What is asymmetry? A body shape that cannot be divided into matching halves. What is radial symmetry? A body arrangement in which body parts are organized around a central axis. What is an example of an animal with radial symmetry? A jellyfish. What is bilateral symmetry? A body arrangement that can be divided into approximately equal left and right halves. What is an example of an animal with bilateral symmetry? A human. What is cephalization? The concentration of sensory organs and nervous tissue toward the front or head region of an animal. What is a body cavity? A fluid-filled space within an animal’s body that can contain and protect internal organs. 3. Major animal groups What are sponges? Simple aquatic animals that lack true tissues and organs. What is the scientific phylum for sponges? Porifera. How do sponges obtain food? They filter small food particles from water flowing through their bodies. What are cnidarians? Aquatic animals with specialized stinging cells. What is the scientific phylum for cnidarians? Cnidaria. What are examples of cnidarians? * Jellyfish * Corals * Sea anemones What are cnidocytes? Specialized stinging cells used for capturing prey and defense. What are arthropods? Animals with segmented bodies, jointed appendages, and an external skeleton. What is an exoskeleton? A hard external structure that supports and protects an animal’s body. What are examples of arthropods? * Insects * Spiders * Crabs * Lobsters What is molting? The process of shedding an old exoskeleton so an arthropod can grow. What are the major animal groups covered in this section? * Porifera: Sponges * Cnidaria: Jellyfish, corals, and sea anemones * Arthropoda: Insects, spiders, and crustaceans Note: Your lecture material did not provide a complete list of characteristics for every animal phylum, so I have not added unsupported details to those sections. Unit 7: Biotechnology 1. Biotechnology basics What is biotechnology? The use of living organisms, cells, or biological processes to develop useful products and technologies. What is genetic engineering? The process of directly changing an organism’s DNA. What is recombinant DNA? DNA made by combining genetic material from different sources. What is a plasmid? A small circular DNA molecule commonly found in bacteria. Why are plasmids useful in biotechnology? They can be used to carry selected genes into cells. 2. PCR What does PCR stand for? Polymerase chain reaction. What is PCR used for? To make many copies of a specific DNA segment. What are the three main stages of PCR? * Denaturation * Annealing * Extension What happens during denaturation? The DNA strands separate. What happens during annealing? Primers attach to complementary DNA sequences. What happens during extension? DNA polymerase adds nucleotides to build new DNA strands. 3. Gel electrophoresis What is gel electrophoresis? A laboratory technique used to separate DNA fragments based on size. How does gel electrophoresis separate DNA? An electric current moves DNA fragments through a gel. Smaller fragments generally travel farther than larger fragments. Why does DNA move toward the positive end of the gel? DNA has a negatively charged phosphate backbone. What can gel electrophoresis help scientists do? Compare DNA fragments and examine DNA samples. 4. Restriction enzymes and cloning What are restriction enzymes? Enzymes that cut DNA at specific sequences. What is DNA cloning? The process of making copies of a DNA segment or gene. What is a GMO? A genetically modified organism whose genetic material has been altered using genetic engineering or other genetic modification methods. What is a transgenic organism? An organism that contains genetic material introduced from another species. What is CRISPR? A gene-editing technology that allows scientists to make targeted changes to DNA. What is gene therapy? A medical approach that aims to treat or prevent disease by changing genetic material in a person’s cells. What is genomics? The study of an organism’s entire genome. What is proteomics? The study of the proteins produced by a cell, tissue, or organism. Practice Questions: Multiple Choice The answer is immediately underneath each question. Try answering before reading it. 1. What is the basic building block of DNA? A. Amino acid B. Nucleotide C. Protein D. Lipid Answer: B. Nucleotide 2. Which base pairs with adenine in DNA? A. Guanine B. Cytosine C. Thymine D. Uracil Answer: C. Thymine 3. Which enzyme separates DNA strands during replication? A. Ligase B. Helicase C. RNA polymerase D. Ribosome Answer: B. Helicase 4. What is the purpose of DNA replication? A. To create proteins B. To make RNA C. To copy DNA before cell division D. To destroy old DNA Answer: C. To copy DNA before cell division 5. What is the process of making mRNA from DNA called? A. Translation B. Transcription C. Replication D. Mutation Answer: B. Transcription 6. Where does translation occur? A. Nucleus B. Ribosome C. Cell membrane D. Golgi apparatus Answer: B. Ribosome 7. What does tRNA carry? A. DNA B. Glucose C. Amino acids D. Lipids Answer: C. Amino acids 8. What is the start codon? A. UAA B. AUG C. UGA D. UAG Answer: B. AUG 9. What is evolution? A. An individual changing during its lifetime B. A population changing genetically over generations C. An organism growing larger D. A cell dividing Answer: B. A population changing genetically over generations 10. Who is associated with evolution by natural selection? A. Gregor Mendel B. Charles Darwin C. Louis Pasteur D. Robert Hooke Answer: B. Charles Darwin 11. What is genetic drift? A. A random change in allele frequencies B. The movement of water C. The creation of proteins D. The copying of DNA Answer: A. A random change in allele frequencies 12. What is gene flow? A. DNA replication B. Movement of alleles between populations C. Protein synthesis D. Natural death Answer: B. Movement of alleles between populations 13. Which equation represents Hardy-Weinberg allele frequencies? A. B. C. D. Answer: B. 14. What are the three domains of life? A. Plants, animals, fungi B. Bacteria, Archaea, Eukarya C. Mammals, reptiles, birds D. Protists, fungi, plants Answer: B. Bacteria, Archaea, Eukarya 15. Which cells do not have a nucleus? A. Eukaryotic B. Prokaryotic C. Animal D. Plant Answer: B. Prokaryotic 16. What is the function of mitochondria? A. Protein storage B. ATP production C. DNA protection D. Photosynthesis Answer: B. ATP production 17. What is the cell wall of fungi made of? A. Cellulose B. Peptidoglycan C. Chitin D. Starch Answer: C. Chitin 18. What tissue transports water in plants? A. Phloem B. Xylem C. Epidermis D. Pollen Answer: B. Xylem 19. What is the function of phloem? A. Transport sugars B. Absorb sunlight C. Produce pollen D. Protect seeds Answer: A. Transport sugars 20. Which plants produce flowers? A. Mosses B. Ferns C. Gymnosperms D. Angiosperms Answer: D. Angiosperms 21. What is the scientific phylum for sponges? A. Cnidaria B. Porifera C. Arthropoda D. Chordata Answer: B. Porifera 22. What do arthropods have? A. Jointed appendages and an exoskeleton B. No body structure C. Only soft bodies D. Chloroplasts Answer: A. Jointed appendages and an exoskeleton 23. What is PCR used for? A. Separating proteins B. Copying DNA C. Producing ATP D. Growing plants Answer: B. Copying DNA 24. What does gel electrophoresis separate DNA by? A. Color B. Shape only C. Fragment size D. Temperature Answer: C. Fragment size 25. What is CRISPR used for? A. Editing DNA B. Separating cells C. Making ATP D. Producing pollen Answer: A. Editing DNA Fill-in-the-Blank Practice 1. DNA stands for __________. Answer: Deoxyribonucleic acid. 2. RNA uses __________ instead of thymine. Answer: Uracil. 3. The enzyme that unzips DNA is __________. Answer: Helicase. 4. The enzyme that connects Okazaki fragments is __________. Answer: DNA ligase. 5. The process of making RNA from DNA is called __________. Answer: Transcription. 6. The process of making proteins from mRNA is called __________. Answer: Translation. 7. A group of three mRNA bases is called a __________. Answer: Codon. 8. The movement of alleles between populations is called __________. Answer: Gene flow. 9. Random changes in allele frequencies are called __________. Answer: Genetic drift. 10. The formation of new species is called __________. Answer: Speciation. 11. The three domains are Bacteria, Archaea, and __________. Answer: Eukarya. 12. The cell wall of fungi contains __________. Answer: Chitin. 13. The plant tissue that transports water is __________. Answer: Xylem. 14. The plant tissue that transports sugars is __________. Answer: Phloem. 15. Flowering plants are called __________. Answer: Angiosperms. 16. The haploid plant stage is called the __________. Answer: Gametophyte. 17. The diploid plant stage is called the __________. Answer: Sporophyte. 18. Sponges belong to the phylum __________. Answer: Porifera. 19. Jellyfish belong to the phylum __________. Answer: Cnidaria. 20. The process used to make many copies of DNA is called __________. Answer: PCR. Important Terms to Memorize Term Meaning DNA Stores genetic information RNA Helps make proteins Nucleotide Building block of DNA and RNA Helicase Separates DNA strands DNA polymerase Builds new DNA DNA ligase Connects DNA fragments Transcription DNA to RNA Translation RNA to protein Codon Three mRNA bases Mutation Change in DNA Evolution Change in inherited traits in a population over generations Natural selection Differences in reproductive success based on inherited traits Genetic drift Random allele frequency change Gene flow Movement of alleles between populations Speciation Formation of new species Taxonomy Naming and classifying organisms Phylogeny Evolutionary relationships Prokaryote Cell without a nucleus Eukaryote Cell with a nucleus Mitochondria ATP production Chloroplast Photosynthesis Xylem Transports water Phloem Transports sugars Gametophyte Haploid plant generation Sporophyte Diploid plant generation Angiosperm Flowering plant Gymnosperm Plant with exposed seeds PCR Makes copies of DNA Gel electrophoresis Separates DNA fragments by size CRISPR Gene-editing technology Final Cram Sheet: What You Should Know Before Your Exam DNA and genetics * DNA: A–T and C–G * RNA: A–U and C–G * Helicase: unzips DNA * DNA polymerase: builds DNA * Ligase: joins DNA fragments * Transcription: DNA → RNA * Translation: RNA → protein * AUG: start codon * UAA, UAG, UGA: stop codons Evolution * Darwin: natural selection * Mutation: changes DNA * Genetic drift: random change * Gene flow: movement between populations * Bottleneck: population becomes much smaller * Founder effect: small group starts a population * Speciation: new species form * Hardy-Weinberg: , Classification and cells * Three domains: Bacteria, Archaea, Eukarya * Prokaryotes: no nucleus * Eukaryotes: have a nucleus * Mitochondria: ATP * Chloroplasts: photosynthesis * Fungi: chitin cell walls * Bacteria: peptidoglycan cell walls Plants * Xylem: water * Phloem: sugar * Gametophyte: haploid * Sporophyte: diploid * Gymnosperms: exposed seeds * Angiosperms: flowers and enclosed seeds * Mosses: nonvascular * Ferns: seedless vascular Animals and biotechnology * Porifera: sponges * Cnidaria: jellyfish, corals, sea anemones * Arthropoda: jointed appendages and exoskeleton * PCR: copies DNA * Gel electrophoresis: separates DNA by size * Restriction enzymes: cut DNA * CRISPR: edits DNA Study tip: Cover the answers with a piece of paper and try to answer every question from memory. Then uncover the answer and check yourself. Focus especially on DNA replication, transcription versus translation, Hardy-Weinberg equations, evolution mechanisms, plant life cycles, and the differences between prokaryotic and eukaryotic cells.