the broad pattern of evolution above the species level
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Protocell
an abiotic precursor of a living. ell that had a membrane-like structure and that maintained an internal chemistry different from that of its surroundings
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Hydrothermal Vents
an area on the deep sea floor where heated water and minerals from Earth’s interior gush into the seawater
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Alkaline Vents
a deep sea hydrothermal vent that releases water that is warm rather than hot and that has a high pH
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Ribozymes
an RNA molecule that functions as an enzyme, such as an intron that catalyzes its own removal during RNA splicing
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Radiometric Dating
a method for determining the absolute age of rocks and fossils, based on the half-life of radioactive isotopes
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Half-Life
amount of time it takes for half of a sample of a radioactive isotope to decay
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Geologic Record
a standard time scale dividing Earth’s history into time periods, grouped into four eons - Hadean, Archaean, Proterozoic, and Phanerozoicand further subdivided into eras, periods, and epochs
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Stromatolites
layered rock that results from the activities of prokaryotes that bind thin films of sediment together
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Endosymbiont Theory
the theory that mitochondria and plastids, including chloroplasts, originated as prokaryotic cells engulfed by a host cell. the engulfed cell and its host cell evolved into a single organism
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Serial Endosymbiosis
a hypothesis for the origin of eukaryotes consisting of a sequence of endosymbiotic events in which mitochondria, chloroplasts, and perhaps other cellular structures were derived from a small prokaryotes that had been engulfed by larger cells
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Cambrian Explosion
a relatively brief time in geologic history when many present-day phyla of animals first appeared in the fossil record. 535-525 million years ago, the emergence of the first large, hard-bodied animals
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Plate Tectonics
the theory that the continents are part of great plates of earth’s crust that float on the hot, underlying portion of the mantle. movements in the mantle cause the continents to move slowly over time
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Pangaea
the supercontinent that formed near the end of the Paleozoic era, when plate movements brought all the landmasses of Earth together
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Mass Extinction
the elimination of a large number of species throughout Earth, the result of global environment changes
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Adaptive Radiations
period of evolutionary change in which groups of organisms form many new species whose adaptions allow them to fill different ecological roles in their communities
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Heterochrony
evolutionary change in the time or rate of an organism’s development
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Paedomorphosis
the retention in an adult organism of the juvenile features of its evolutionary ancestors
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Homeotic Genes
any of the master regulatory genes that control placement and spatial organization of body parts in animals, plants, and fungi by controlling the development fate of groups of cells
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Phylogeny
the evolution of history of a species or group of related species
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Systematics
a scientific discipline focused on classifying organisms and determining their evolutionary relationships
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Taxonomy
a scientific discipline concerned with naming and classifying the diverse forms of life
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Binomial
the two-part latinized format for naming a species, consisting of the genus and specific epithet
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Linean Classification
Domain → Kingdom → Phylum → Class → Order → Family → Genus → Species
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Taxon
a named taxonomic unit at any given level of classification
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Phylogenetic tree
a branching diagram that represents a hypothesis about the evolutionary history of a group of organisms
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Phylocode
proposed system of classification of organisms based on evolutionary relationships: only groups that include a common ancestor and all of its descendants are named
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Branch Points
the representation on a phylogenetic tree of the divergence of two or more taxa from a common ancestor
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Rooted
describing a phylogenetic tree that contains a branch point representing the most recent common ancestor of all taxa in the tree
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Basal taxon
in a specified group of organisms, a taxon whose evolutionary lineage diverged early in the history of the group
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Polytomy
in a phylogenetic tree, a branch point from which more than two descendant taxa emerge. a polytomy indicates that the evolutionary relationships between the descendant taxa are not yet clear
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Analogy
similarity between two species that is due to convergent evolution rather than to descent from a common ancestor with the same trait
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Homoplasies
a similar structure or molecular sequence that has evolved independently in two species
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Molecular Systematics
a scientific discipline that uses nucleic acids or other molecules to infer evolutionary relationships between different species
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Cladistics
an approach to systematics in which organisms are placed into groups called clades based primarily on common descent
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Clades
a group of species that includes an ancestral species and all of its descendants
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Monophyletic
pertaining to a group of taxa that consists of a common ancestor and all of its descendants. a monophyletic taxon is equivalent to a clade
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Paraphyletic
pertaining to a group of taza that consists of a common ancestor and some, but not all, of its descendants
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Polyphyletic
pertaining to a group of taxa derived from two or more different ancestors
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Shared Ancestral Character
a character, shared by members of a particular clade, that originated in an ancestor that is not a member of taht clade
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Shared Derived Character
an evolutionary novelty that is unique to a particular clade
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Outgroup
a species or group of species from an evolutionary lineage that is know to have diverged before the lineage that contains the group of species being studied. an outgroup is selected so that its members are closely to the group of species being studied, but not as closely related as any study-group members are to each other
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Ingroup
a species or group of species whose evolutionary relationships we seek to determine
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Maximum Parsimony
a principle that states that when considering multiple explanations for an observation, one should first investigate the simplest explanation that is consistent with the facts
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Maximum Likelihood
as applied to molecular systematics, a principle that states that when considering multiple phylogenetic hypotheses, one should take into account the hypothesis that reflects the most likely sequence of evolutionary events, given certain rules about how DNA changes ove time
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Orthologous Genes
homologous genes that are found in different species because of speciation
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Paralogous Genes
homologous genes that are found in the same genome as a result of gene duplication
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Molecular Clock
a method for estimating the time required for a given amount of evolutionary change, based on the observation that some regions of genomes evolve at a constant rate
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Neutral Theory
the hypothesis that much evolutionary change in genes and proteins has no effect on fitness and therefore is not influenced by natural selection
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Horizontal Gene Transfer
the transfer of genes from one genome to another through mechanisms such as transposable elements, plasmid exchange, viral activity, and perhaps fusion of different organisms
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Peptidoglycan
a network of sugar polymers cross-linked by polypeptides
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Gram Stain
used to classify bacteria by cell wall composition
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Gram-Positive Bacteria
simpler walls with a large amount of peptidoglycan (stains purple)
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Gram-Negative Bacteria
have less peptidoglycan and an outer membrane that can be toxic (stains red)
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Capsule
the cell wall of many prokaryotes is surrounded by a sticky layer of polysaccharide or proteins
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Endospores
a way of withstanding harsh conditions, certain bacteria develop resistant cell called this
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Fimbriae
hairlike appendages that stick to substrates
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Pili (sex-pili)
appendages that pull two cells together prior to DNA transfer from one cell to another
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Taxis
the ability to move toward or away from a stimulus
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Nucleoid Region
chromosome is located here
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Plasmids
some species of bacteria also have smaller rings of independently replicating DNA called this
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Three factors that contribute to genetic diversity
the combining of DNA from two sources, contributes to diversity
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Transformation
a prokaryotic cell can take up and incorporate foreign DNA from the surrounding environment
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Transduction
the movement of genes between bacteria by bacteriophages (viruses that infect bacteria)
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Conjugation
the process where genetic material is transferred between prokaryotic cells
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F Factor
piece of DNA that is required for the production of pili
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F Plasmid
cells containing this functions as DNA donors during conjugation
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R Plasmid
carry genes for antibiotic resistance
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Phototrophs
obtain energy from light
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Chemotrophs
obtain energy from chemicals
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Autotrophs
require CO2 as a carbon source
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Heterotrophs
require an organic nutrient to make organic compounds
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Obligate Aerobes
require O2 for cellular respiration
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Obligate Anaerobes
are poisoned by O2 and use fermentation or anaerobic respiration
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Facultative Anaerobes
can survive with or without O2
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Nitrogen Fixation
some prokaryotes convert atmospheric nitrogen (N2) to ammonia (NH3)
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Heterocysts
nitrogen-fixing cells
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Extreme halophiles
live in highly saline environments
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Extreme Thermophiles
thrive in very hot environments
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methanogens
live in swamps and marshes and produce methane as a waste product
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Decomposers
breaking down dead organisms and waste products
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Symbiosis
an ecological relationship in which two species live in close contact. a large host and a smaller symbiont
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Mutualism
both symbiotic organisms benefit
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Commensalism
one organism benefits while neither harming nor helping the other in any significant way
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Parasitism
an organism called a parasite harms but does not kill its host
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Pathogens
parasites that cause disease
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Exotozins
secreted and cause disease even if the prokaryotes that produce them are not present
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Endotoxins
released only when bacteria die and their cell walls break down
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Algae (alga)
a photosynthetic, plantlike protist
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Alternation of Generations
a life cycle in which there is both a multicellular diploid form, the sporophyte, and a multicellular haploid form, the gametophyte; characteristic of plants
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Alveolata
a protistan clade that includes dinoflagellates, apicomplexans, and the ciliates. have small membrane-bounded cavities called alveoli under their cell surfaces. function is unknown
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Amoeba
a type of protist characterized by great flexibility and the presence of pseudopodia
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Apicomplexan
one of a group of parasitic protozoans, some of which cause human diseases
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Blades
a leaflike structure of a seaweed that provides most of the surface area for photosynthesis
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Brown Algea
one of a group of marine, multicellular, autotrophic protists, the most common type of seaweed. include the kelps
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Cellular Slime Mold
a type of protist that has unicellular amoeboid cells and multicellular reproductive bodies in its life cycle
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Ciliate
a type of protozoan that moves by means of cilia
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Conjugation in bacteria
the direct transfer of DNA between two cells that are temporarily joined
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Diatom
a unicellular photosynthetic alga with a unique, glassy cell wall containing silica