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part of the retina that receives light from the center of the visual field
fovea
most sensitive part of the retina
fovea
part of the retina where axons from ganglion cells leave the retina here to form the optic nerve; no photoreceptors here
optic disk (blind spot)
only cell in the retina that fires action potentials
ganglion cells
parts of retina that release neurotransmitter proportional to their level of depolarization (a graded response)
photoreceptors and bipolar cells
2 types of photoreceptors
rods and cones
photoreceptor that enables vision in low-light conditions
rods
photoreceptor that enables high-acuity and color vision in good lighting conditions
cones
where is the highest density of cones?
fovea
where is the highest density of rods?
retinal periphery
in what conditions do sodium channels open?
dark
in what conditions do sodium channels close?
light
when are photoreceptors depolarized?
in dark
when do photoreceptors release glutamate?
in dark
when do photoreceptors hyperpolarize?
in light
when do photoreceptors inhibit glutamate release?
in light
enables contrast sensitivity and edge detection
center-surround receptive fields
cell that receives direct input from photoreceptors
bipolar cells
bipolar cells that respond well to light in the center of the receptive field (not in the surround)
ON-center
bipolar cells that respond well to light in the surround of the receptive field (not in the center)
OFF-center
cells that provide output from retina
ganglion cells
2 main types of receptive fields
ON-center and OFF-center
ganglion cell that responds with either a) the onset of small bright light in center of receptive field or b) the onset of annulus of light in surround of receptive field
ON-center
what are ganglion cells sensitive to?
contrast (adjacent light and dark areas)
3 types of cones
short (blue), medium (green), long (red)
what color corresponds to short wavelength cones?
blue
what color corresponds to medium wavelength cones?
green
what color corresponds to long wavelength cones?
red
theory that states there are 3 cone types at each point on the retina, white = all 3 cones active, colors from different combinations
trichromacy
theory that states that red and green are opponent colors and blue and yellow are opponent colors
color-opponency
what are the 3 channels that carry info from retina to V1?
2 chromatic channels and 1 luminance channel
3 types of ganglion cells
parasol, midget, small bistratified
which ganglion cell contributes to luminance channel?
parasol
which ganglion cell contributes to red-green channel?
midget
which ganglion cell contributes to blue-yellow channel?
small bistratified
in what layers of the LGN are magnocellular neurons in?
1-2
in what layers are parvocellular neurons in?
3-6
in what layers are koniocellular neurons in?
K1-K6
what color cells are in parvocellular layers?
red/green
what color cells are in koniocellular layers?
blue/yellow
what color cells are in magnocellular layers?
black/white
ganglion cell that processes form and color info
parvocellular
ganglion cell that processes color info
koniocellular
ganglion cell that processes motion and depth info
magnocellular
what layers in V1 do magnocellular neuron pathways project to?
sublayer 4Cα
what layers in V1 do parvocellular pathways project to?
sublayer 4Cβ
what layers do koniocellular pathways project to?
layers 1-3
cell that has distinct ON and OFF regions (yields orientation selectivity)
simple cell
cell that has no distinct ON and OFF regions
complex cell
high amounts of cytochrome oxidase that occur at regular intervals in V1; are thought to contribute to color processing
blobs
what are higher-order visual cortical cells selective for?
complex objects or motion direction
audible sound range
0-120 db
rapid pressure fluctuations in a medium
sound
measures the magnitude of pressure fluctuations
sound pressure level
human hearing range
20-20,000 Hz
sound travel speed
340 m/s
3 main cues to localize sound
interaural time difference, interaural level difference, head-related transfer function
difference in time taken for a sound to reach each ear, binaural cue
interaural time difference (ITD)
difference between the sound pressure level at each ear, binaural cue
interaural level difference (ILD)
pinna, head and torso influence the sound before it reaches the inner ear, spectral cue
head-related transfer function (HRTF)
when do we mainly use ITD?
low sound frequencies
when do we mainly use ILD?
high sound frequencies
what are ITD and ILD mainly useful as?
horizontal location cues
sound moves:
tympanic membrane
ossicles move with:
tympanic membrane
malleus, incus, stapes
ossicles
ossicles move:
oval window
how do ossicles act as an amplifier?
convert air movement into cochlear fluid movement
parts of inner ear
cochlear fluids, basilar membrane, inner hair cells, outer hair cells
low K+ cochlear fluid in scala tympani and scala vestibuli
perilymph
high K+ cochlear fluid in scala media
endolymph
what part of the inner ear moves up and down with sound?
basilar membrane
what part of the inner ear transmits info to the brain?
inner hair cells
what part of the inner ear amplifies movement of basilar membrane?
outer hair cells
where is the maximal movement of the cochlear membrane?
at the base for high-frequency sounds and at the apex for low-frequency sounds
when do hair cells depolarize?
when stereocilia deflect towards kinocilium
when do hair cells hyperpolarize?
when stereocilia deflect away from kinocilium
how does the cochlea send info?
via auditory nerve
neurons fire action potentials at a particular phase of the sound wave
phase-locking
when does phase-locking occur?
at low frequencies (<4 kHz)
how is the auditory nerve arranged?
tonotopically (carries info to cochlear nucleus)
what is the first site to receive input from both ears?
superior olive
relays info to olive, preserves response timing of auditory nerve
ventral cochlear nucleus
complex spectral analysis, projects directly to inferior colliculus
dorsal cochlear nucleus
integrates spatial and spectral analysis
inferior colliculus
earliest area sensitive to binaural cues
superior olive
where are ITDs encoded?
medial superior olive
where are ILDs encoded?
lateral superior olive
occupies dorsal and lateral superior temporal gyrus
auditory cortex
auditory cortex regions
core, belt, parabelt
region that includes primary auditory cortex
core
region that surrounds the core region, particularly anterolaterally
belt
region that is posterolateral superior temporal gyrus
parabelt
3 areas of auditory cortex core
primary auditory cortex, rostral field, rostrotemporal field
what sounds does the core respond to?
pure tones
what sounds does the belt respond to?
band-passed noise
what sounds does the parabelt respond to?
complex sounds
what does the auditory ventral "what" pathway process?
auditory objects
what does the auditory dorsal "where/how" pathway process?
sound location
what does the posterior superior temporal gyrus represent?
fast varying speech sounds