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Ecology
The scientific study of the relationships between organisms and their environment
Ecological hierarchy
Individual, population, community, ecosystem, landscape, biome, biosphere
Abiotic
Physical and chemical conditions (abiotic factors) include;
• temperature
• moisture
• concentration of gases (O2 and CO2)
• light intensity
Biotic
Interactions with other organisms (biotic factors) include;
• mating
• predator and prey
Individual
single organism
• Smallest and simplest unit of ecology
• Sense and responds to its environment
• Reproduces -> passes on genetic information the next
generation -> contributes to future populations,
communities, ecosystems
• Birth and death are one-time events for each individual
Population
group of individuals of the same
species, found in a defined area
• Instead of discrete events, birth and death are
continuous
• measured as rates, such as births/year
Ecosystem
communities + habitats (biotic AND
abiotic components)
• emphasis shifts from species to the flows of energy and
nutrients through both biotic and abiotic components
of the ecosystem
Community
different populations of living
organisms within a specific area (interacting biotic
components)
• Focus shifts to relative abundance of species within the
community and
• How interactions among species affect birth and death
rates
Landscape
a patchwork of multiple interacting
ecosystems in which spatial patterns are
important
Biome
large scale region with similar climatic
conditions (tundra, tropical rainforest)
• focus on the distribution of different types of ecosystems
Biosphere
global zone encompassing all life on
Earth
• emphasis on the linkages between ecosystems and
other components of Earth, such as the atmosphere
• How are nutrients such as carbon exchanged between
the atmosphere and terrestrial ecosystems
Weather
can influence reproduction in plant populations, where ______ patterns are influenced by solar radiation with heat coming from shorter wavelengths- higher energy. The average net radiation = zero, half absorbed and half reflected
Insolation
Amount of solar radiation reaching surface, varies w/ latitude; shorter distance around the equator
Latitude
Short at equator, long at poles, more insolation at equator
Diurnal cycle
The hours of daylight and dark that vary with seasons except at the equator, which has 12 hrs day and 12 hrs night throughout the year
Climate
can influence succession from prairie to a forest
Bergmann's rule: Cold-climate animals are often larger than warm-climate animals, mostly applies to birds & mammals
Bergmann’s rule
Cold-climate animals are often larger than warm-climate animals, mostly applies to birds & mammals
Natural selection
differential success (survival
and reproduction) of individuals within a
population
• interactions with the environment are the
selective agents
Requirements:
1. Variation in traits must be heritable
2. Variation leads to differences in survival and
reproduction (fitness) among individuals in the
population
Changes in population leads to evolution over
time
Fitness
the proportionate contribution made by an individual to future
generations relative to other individuals in the same population
• Traits that confer higher fitness are more frequent in the next generation
Greater fitness → Higher Rates of Survival + Reproduction = More Offspring
Adaption
any heritable behavioral,
morphological, or physiological trait that
has evolved through the process of
natural selection
Phenotypic plasticity
the ability of
one genotype to give rise to different
phenotypes under different
environmental conditions, leads to increased fitness
Developmental plasticity
phenotypic changes cannot be reversed
Acclimation
phenotypic plasticity in
response to current environmental
conditions that is reversible
• fish have upper and lower limits to
temperatures they can tolerate
• these limits change as water
temperature changes with the
seasons
Subpopulations
local populations of interbreeding individuals linked by movement of individuals
Selective agent
environmental cause of
fitness differences among organisms with
different phenotypes
Directional selection
Occurs when the phenotypes
at one end of the distribution
have higher fitness than the
mean phenotype; only one extreme high/low favored
Disruptive selection
Occurs when the phenotypes
at both ends of the distribution
have higher fitness than the
mean phenotype; both high/low favored
Stabilizing selection
Occurs when the mean phenotype
has higher fitness than the
phenotypes at either end of the
distribution; average favored
Genetic drift
change in allele frequencies as a result of random chance
• Changes are random, not fitness based
Gene flow
the movement of genetic information among populations
• Reduces variation among populations by keeping allele frequencies more similar
Assortative mating
individuals in a population choose mates based on
their phenotype, which reflects their genotype
Positive assortative mating
mates are phenotypically more similar to each
other than expected by random chance
• Increases homozygosity
Negative assortative mating
mates are phenotypically less similar to each
other than expected by random chance
• Increases heterozygosity
Inbreeding
individuals mate with other
members of the population who are more
closely related to them than expected by
random chance
• This increases homozygosity at all genes
Inbreeding depression
offspring are more
likely to homozygous for harmful recessive
alleles that can lead to reductions in fertility,
vigor, fitness and even death
Cline
a measurable change in a phenotypic character or characters
over a geographic region or a gradient in genotypic frequency
Step clines
abrupt changes in local environments and in the phenotype and
genotype of the organisms
Ecotypes
population adapted to its unique local environmental conditions
Trade-off
Constrains the evolution of adaptions with the constraints being physiological, energetic, and environmental
Adaptive radiation
multiple species within a
single lineage that exploit
different features of the
environment
Light dependent photosynthesis
• Chlorophyll absorbs light
energy
• Light energy excites electrons,
splits H2O molecules
• Light energy is Invested into
ATP and NADPH
Light independent photosynthesis
• Carbon Fixation occurs during
the Calvin Cycle (C3 Cycle)
• ATP and NADPH are used with
CO2 to make glucose
Stomata
openings on the
leaf surface that allow CO2
to enter
Transpiration
when the stomata are
open, water vapor in the leaf diffuses out
Turgor pressure
the force exerted
outward on a cell wall by the water
inside the cell
C3
plants in areas without water
limitation generally use __
photosynthesis = cool moist
environments
• Stomata remains open -> inefficient in
hot, dry conditions
• CO2 is fixed in a single step within
mesophyll cells, directly forming a
three-carbon compound via the Calvin
Cycle
C4
plants typically in arid, salty, tropical or
subtropical environments
• Carbon fixation is a two-step process with spatial
separation in the leaf
1. CO₂ fixed into a four-carbon compound in
mesophyll cells
2. Released in bundle sheath cells for the Calvin
Cycle to use
• Effectively concentrates CO₂ = minimizes
photorespiration and enhance efficiency in hot
environments -> stomata opens less = less water loss
CAM
adaption to deserts, extremely low
rainfall
• Carbon fixation occurs at night- stomata stay
closed during the day to prevent water loss
1. Night- stomata is open, CO2 convert to malic
acid
• large amounts of malic acid accumulate in
mesophyll cells
2. Day- stomata is closed, malic acid converted
to CO2
• C3 pathway used to fix CO2 and produce sugars
Dry environment
Adaptions to a ____________________ include increased root production, reduction in leaf area, changes in leaf size and shape, dropping leaves before dry season
Cold environment
Adaptions to a ____________________ include frost hardening, producing compounds that allow leaves to survive freezing temperatures, and some species being winter deciduous
Aquatic environment
Adaptions to a _______________ include floating broad leaves with a waxy cuticle, reproductive parts extending out of the water, highly dissected flexible leaves and stems, reduced roots
Low nutrients
Plant adaptations to _______________ include being carnivorous, having mycorrhizal associations, nitrogen fixers
Animal
___________ adaptations to the environment include gas exchange and acquisition of energy
Herbivory
Plant adaptations to deter ________ include thorns/spines, prickles/hairs, tough leaves/bark, toxins and poisons, bitter compounds, sticky resins
Grazer
Type of herbivore that eats mainly leaves, especially grasses
Browser
Type of herbivore that eats woody material
Granivore
Type of herbivore that eats seeds
Frugivore
Type of herbivore that eats fruit
Herbivore
_____________ adaptations to the environment include having specialized mouthparts, long digestive tracks with symbiotic microorganisms to break down cellulose, and chemical receptors in the nose/mouth to discourage eating certain foods
Carnivore
______________ adaptations to the environment include having strong jaws, sharp teeth and claws, hunting strategies, keen senses, and short digestive tracts with high stomach acidity
Omnivore
___________ adaptations include having sharp front teeth and molar back teeth, foraging, medium length intestines to digest a variety of foods, acute sensory systems
Ectothermy
process of maintaining body temperature through exchange
of thermal energy with the surrounding environment
Endothermy
process of maintaining body temperature through internally
generated metabolic heat
Endothermic homeotherms
bird and mammals
• Can maintain a constant body temperature
through metabolic processes = high metabolic
rate
Ectothermic poikilotherms
typically all other
animals that aren't birds and mammals
• Body temperature varies with the environment
= low metabolic rate
Poikilotherm
________________ adaptations to changing temps include seeking out appropriate microclimates (finding sun/shade), undergo temperature acclimation, change conduction of heat, tolerance to freezing
Homeotherm
___________ adaptations to changing temps include insulation, shivering, brown fat, and decrease heat by evaporative cooling where moisture evaporates from the skin, losing heat
Temporal heterotherms
have characteristics of endotherms at some times and
characteristics of ecotherms at other times
• May be daily or seasonally or only in certain situations
Torpor
the dropping of the body temperature to approximately ambient temperature for part of a day
Metapopulations
collection of local
subpopulations linked through dispersal
or other means of genetic exchange
Abundance
Number of individuals in a population
Population density
The number of individuals per unit area of a population
Population size
Abundance is found by; population density * the area occupied
Absolute density
Complete count of individuals/area; 25 gopher tortoises/hectare
Relative density
Proportion of species relative to others in a community; 25% sunflowers/m2
Quadrat sampling
Sampling area of a given shape, count individuals in quadrats, doesn't work well on motile organisms
Transects
Sample along a line or path, record organisms that touch/are close to the line at set intervals
Mark-recapture
Capture, mark, release, recapture; good for mobile organisms
Lincoln-Petersen
Most common method for mark-recapture; N=nM/R
Two sampling periods- one for marking, one for recapturing
N = Number of animals in the population
M = Number of animals marked on the first visit
n = Number of animals captured on the second visit
R = Number of recaptured animals that were marked
Traps
live animal traps, light traps, pitfall traps, plankton nets
• number of individuals caught per unit time per trap
Catch per unit fishing effort
Catch (number or weight of fish) over the Unit of effort (time, gear, distance)
Vocalization frequency
Good for frogs/bats that may not be visible during other population surveys, calls/unit time
Natality
number of births
related to population size
• Example- number of offspring/1000
indiv./unit time
• includes hatching, germination, fission
Fecundity
number of
offspring/female/unit time
Population growth
To study this, we need to know the starting population size (N0), the net reproductive rate (R0, how many females produced by an average female), are generations discrete or overlapping, if the population is closed or open, time period of measurement or number of generations, the intrinsic rate of increase (R) per capita ate of growth of population w/ stable age distribution, environmental influences
Net reproductive rate
R0 = sum of lx * bx
lx = survivorship, the number of individuals surviving to a given age (x) as a proportion of
the original cohort size, nx/n0
bx = fecundity, the mean number of females born to each female in an age group
Life table
an age-specific account of mortality to see the patterns of mortality and survivorship within populations.
Cohort
Followed by life tables, are group of individuals in a population born in the same period of time
Geometric growth model
predicts changes in population size in discrete intervals where birth and
death are not continuous processes; unlimited resources where the growth rate isn't influenced by population size. step-wise J-shaped curve, discrete generations
Exponential growth model
continuous, unlimited growth,
overlapping generations, favorable environments with unlimited resources and no competition, smooth J-shape curve, overlapping generations
Logistic growth
found where environmental limits exist, growth
rate is influenced by population size, smooth s-shape curve, overlapping generations
Geometric
Discrete time, N(t)=N(0)*lambda^t, where lambda is the finite rate of increase
Exponential
Continuous time, N(t)=N(0)e^rt where r is the intrinsic rate of increase
Stochasticity
random, unpredictable fluctuations that influence population dynamics
Altricial
Young are born helpless, less energy before birth but more care after birth, requires parental care to survive; birds and most mammals
Precocial
Young born in advanced stage of development, can forage independently shortly after birth, requiring less parental care; ducks, geese, horses, giraffes, whales, dolphins, hares
Iteroparous
Organisms reproduce more than once in their lifetime
Semelparous
Organisms reproduce only once in their lifetime
Outcrossing
Plant mating system where cross-fertilization happens between two individuals
Autogamy
Plant mating system where self-fertilization happens