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What is evolution?
a change in gene frequencies over time
What is macroevolution?
changes in allele frequency within the entire biota over long periods of time
What is phylogeny?
the evolutionary history of a group
What is a phylogenetic tree?
a graphical summary of the evolutionary history
What does the phylogenetic tree illustrate?
the sequence of speciation events, which taxa are more closely or distantly related, and sometimes the timing of branching events
What are synapomorphies?
shared, derived traits that evolved in a common ancestor and are present in all or most of its descendants
What is homoplasy?
when a trait is shared by different organisms but didn’t come from a common ancestor
What is the principle of parsimony?
the preferred tree is the one that minimizes the total amount of evolutionary change that has occured
What is the principle of parsimony used for?
when characters give conflicting information about phylogeny, it can be used to decide which tree most closely reflects evolutionary history
What are the three phylogenetic groups?
monophyletic, polyphyletic, and paraphyletic
What is a monophyletic group?
contains a common ancestor and all of its descendants
What is another name for a molophyletic group?
a clade
What is another name for a clade?
a monophyletic group
What is an example of a monophyletic group?
cnidarians (corals, jellyfish, anemones)
What is a paraphyletic group?
consists of an ancestor and some, but not all, of its desendants
What is an example of a paraphyletic group?
fish, includes sharks, rays, bony fish, but excludes tetrapods
What is a polyphyletic group?
consists of organisms that are grouped together but don’t share their most common ancestor within that group
What is an example of a polyphyletic group?
seaweeds
What are the patterns of macroevolution?
stasis, character change, lineage-splitting (speciation), and extinction
What is stasis?
lineages that don’t change much for a long time
What are organisms in stasis called?
living fossils
What are examples of stasis organisms?
coelacanths and horseshoe crabs
What is character change?
lineages can change quickly or slowly, in a single direction, or can reverse
What is an example in character change?
rib number in trilobite lineages
What is lineage-splitting (speciation)?
lineages splitting into separate species
What is extinction?
can be a frequent or rare event within a lineage, or it can occur simultaneously across many lineages
What are the mechanisms of macroevolution?
genetic drift and natural selection
What is genetic drift?
random changes in allele frequencies due to chance
What is natural selection?
non-random change in allele frequencies because some traits increase survival or reproduction
What are the active research questions in evolutionary theory?
does evolution tend to proceed slowly and steadily or in quick jumps and why are some clades very diverse and some unusually sparse
What is a gradualism model of evolution?
expectation of fossil record to preserve many traditional forms, documenting a slow, continuous transformational from ancestral to descendant species over long time
What is rapid burst evolution?
expect fossil records to show sudden morphological changes over short time periods, with relatively few transitional forms linking ancestral and descendant species
What does clade diversity depend on?
speciation rate and extinction rate
In what rate does a clade grow?
speciation higher than extinction
How does the clade move when speciation is more than evolution?
the clade grows
What is the rate when a clade shrinks?
speciation is less than extinction
How does the clade move when evolution is higher than speciation?
the clade shrinks
What rate does the clade stay stable at?
speciation equals extinction
How does the clade move when speciation equals extinction?
the clade stays stable
What traits increase speciation?
small body size and limited dispersal
What traits reduce extinction?
large geographic range/high dispersal potential and broad environmental tolerance
What is evolutionary potential?
the capacity of a population to evolve in response to environmental change
What are the levels of taxonomic classification?
domain, kingdom, phylum, class, order, family, genus, and species
What are viruses?
parasites that develop and reproduce only when infecting a living prokaryotic or eukaryotic cell
What do viruses consist of?
a small amount of DNA or RNA protected by an outer protein coat?
What is the outer protein coat in viruses?
capsid
What is capsid?
the outer protein coat in viruses
What are the smallest microbes?
viruses
How big are viruses?
20-200 nm
What are the building blocks of life?
carbon, nitrogen, and phosphorous
What is carbon?
the chemical backbone of living material
What is nitrogen?
a limiting nutrient of primary production
What is phosphorous?
an essential element of nucleic acids and lipids
Where does DOM come from?
half of all organic matter produced by primary producers
What is the microbial loop?
the major energy pathway where DOM isn’t lost from the system
What uses the dissolved organic matter?
bacteria and archaea
What eats bacteria and archaea?
tiny protozoans (protists)
What eats the protozoans?
zooplankton
What is the largest source of DOM?
virus activity
What is cell lysing?
viruses causing infected cells to burst and release the cells’ contents
What are prokaryotes?
the smallest and simplest independently living things on Earth
What do prokaryotes include?
eubacteria (bacteria and cyanobacteria) and archaea
What are prokaryotic cells enclosed by?
a protective outer cell wall
What is the cell wall in prokaryotes?
the outer protection
What do prokaryotes lack?
a nucleus and membrane bound organelles
What is photoautotrophy?
using photosynthetic pigments to trap light energy to manufacture organic compounds
What is chemoautotrophy?
deriving energy from chemical compounds to manufacture organic compounds
How big are bacteria?
about 1 micron long
What is ubiquity?
found everywhere
What is the ecological role of bacteria
important part of microbial loop, consuming DOM released when viruses lyse cells, passing energy up to protozoans
What else in cyanobacteria called?
blue-green algae
What group are cyanobacteria in?
photosynthetic bacteria
What is the most abundant photosynthetic organism in the ocean?
prochlorococcus
What is among the first photosynthetic organisms on Earth?
cyanobacteria
What are stromatolites?
calcareous mounds formed by cyanobacteria
What is the ecological role of cyanobacteria?
accumulation of oxygen in atmosphere
What is symbiotic bacteria?
bacteria evolved to live in close association with other marine organisms
What is mutualism?
interactions where both parties benefit
Where are bacteria sheltered in other organisms?
in special tissues or organs
What are archaea?
prokaryotic microorganisms once thought to be bacteria but now recognized as being more closely related to eukaryotes
What where archaea first known as?
extermophiles
What is extremophily?
living in extreme environments
What is the ecological role of archaea?
not well understood
What are eukaryotes?
cells that have membrane-bound nuclei
What are protists or protozoans?
unicellular eukaryotic organisms
What is autotrophy?
plank-like
What is heterotrophy?
animal-like
What are the plank-like protists?
diatoms, dinoflagellates, and coccolithophores
What are diatoms?
cells enclosed by cell wall of silica and form a glassy shell or frustule
What are the small holes in frustule for?
allowing water and nutrients in and out
What is the ecological role of diatoms?
important primary producers in temperate and polar zones?
What percentage of global photosynthetic fixation of carbon do diatoms account for?
20%
What are dinoflagellates?
cell wall armored with plates of cellulose with spines, pores, etc.
What do dinoflagellates have?
two flagella
Where are the flagella of dinoflagellates located?
around the groove of middle of cell and trailing
What is the purpose of the middle flagella?
forward motion and spinning
What is the purpose of the trailing flagella?
propulsion
What is the ecological role of dinoflagellates?
important primary producers in tropical and subtropical zones
What are dinoflagellates often responsible for?
harmful algal blooms
What are coccolithophores?
flagellated, spherical cells, covered with coccoliths plates made of calcium carbonate