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Catastrophism
Idea that changes to Earth’s crust happen because of sudden, violent events such as natural disasters and not gradual change, believed until the 1700s
Uniformitarianism
Theory that changes to Earth’s crust happen due to continuous and uniform process, founded by James Hutton in 1785
Continental Drift
Idea that the continents were once a whole mass of land called Pangea and have since drifted over geologic time, proposed by Alfred Wegener in 1912, Pangea is said to have broken up 200 mya.
Plate tectonics
Provided an explanation for Wegener’s lack of mechanism in his hypothesis, unifying model which accounts for origin of continents and oceans, mountain ranges, earthquakes, etc.Â
Internal structure of Earth
Has a thermal gradient (hotter in the center)
Inner core: solid due to high pressure
Outer core: molten iron
Mantle: mostly solid rock of iron and magnesium silicate (flows slowly)
Crust: mostly granite (continents) and basalt (oceans)
Lithosphere: crust and upper mantle which is broken into 20 plates that can move over plastic mantle
Fossils and genetic evidence for Pangea
There are matching fossil records that support Pangea, current distribution of taxa also supports. Example: Araucaria is a plant that is only in southern hemisphere
Age of Earth
4.5 billion years old (Ga)
Stromatolites
Earliest microfossils from 3.5 Ga
Cyanobacteria
Earliest evidence from 2.7-2.1 Ga and are disputed fossils
Algae-like plants
Showed up at 1 GaEm
Embryophytes
Land plants, appeared in mid-Ordovician period (470 mya)EM
Embryo
Zygote protected by female gametophyte and is essential for survival on land
Alternation of generations
Facilitated by:
Haploid gametophyte (makes gametes)
Diploid sporophyte (makes spores)
Sporopollenin and cuticle
Dessication-resistant compounds, prevents plant from becoming extremely dried out
Carpet bombing
Spores are major agents of dispersal, many spores released at once
Vascular plants
Seedless, appeared in late Silurian/early Devonian (about 420 mya)
Seed plants
Also vascular, appeared during Devonian (about 380 mya), seeds are agent of dispersal, characterized by:
Heterospory - producing 2 kinds of spores
Megaspores - retained by sporophyte (protects embryo)
Microspores - travels to fertilize megaspore
Domestication syndrome
Selection by humans which has led to plants characterized by a set of phenotypic traits not adaptive in the wild, dependent on humans for survival
Landraces
Local variety of a domesticated plant, differ in adaptations to environments as well as culinary uses, Example: different kinds of maiz
Nikolai Vavilov
One of first to note lower resilience of domesticated plants compared with wild relatives
Proposed interbreeding
Collected germplasm of domesticated + wild
Identified 8 centers of origin for domestication
Centers of Origin
China (rice, soybeans, peach)
A) Indo-Malay (banana, coconut) B)Indo burma (mango, eggplant)
Central Asia (lentils, apples)
Fertile Crescent (oats, wheat, dates)
Mediterranean (olives, cabbage, mint)
Ethiopia (coffee, okra, sesame)
Mesoamerica (corn, chocolate, tomato)
A) Andes (potato, lima beans, corn) B) Chile (some potatoes strawberries) Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â Â C) Chaco-Pantal (cashew, rubber, pineapple)
Population
A group of freely interbreeding individuals of the same species in the same habitat, species can have multiple populations
Annual
Completes life cycle (seed to seed) within 1 year or less
Biennial
Tends to live for 2 years, 1st year grows roots and leaves, 2nd year long stems and flowers, dies after setting seed
Perennial
Lives several years and usually flowers several times, can be woody or survive underground (tubers)
Semelparous
Reproduces only once in its lifetime, includes annuals, biennials, and some perennials
Iteroparous
Reproduces many times during its life, includes most perennial plants
Hermaphroditism
Flowers with both male and female gametes
Monoecious
Plants with both male and female flowers
Dioecious
Plants with either male or female flowers
r selection
Many offspring, low investment, short life, most die before maturity, low individuality, etc.
K selection
Few offspring, long life, high investment, high individuality, etc.
Macroenvironment
Elevation, latitude, temperature, rainfall, etc.Â
Microenvironment
Localized changes in light availability, moisture, nutrients, etc.
Taxonomic groupings that reflect geological and evolutionary history
Holarctic
South American
African
Indo-Pacific
Australian
Biomes/types of vegetation
Tundra: Small-sized plants adapted to short seasons
Taiga: Conifer forests
Deciduous forests: Broad-leaf temperate forests
Grasslands: Praires, steppes, savannah, pampas
Shrublands: Chaparral, scrub, etc.
Desert: Plants are further apart than in shrublands
Tropical forests: human and warm, peak diversity
Biotic factors
Result from interactions with other organisms, Example: herbivory, infection, symbiosis, etc.
Abiotic factors
Includes parameters (temperature, pH, humidity) and resources (light, water, minerals) that determine growth as well as others that can add stress (pollution)
Limiting factors
Can limit plant processes, can be caused by one environmental factor at suboptimal level
Stress
Occurs when essential factors (light, water, oxygen, nutrients) are at inappropriate levels
Environmental noxa
Factors that trigger stress at ANY concentration (heavy metals, pollutants, etc.)
State of latent life
A state which plants reach when intensity of stress factors is inappropriate where minimal life processes occur
Dormancy
Adapted latent life stage triggered by environmental cute, involves physiological changes
Avoidance strategy
Seeds and spores allow annual/ephemeral plants to avoid moments of highest environmental stress
Mesophiles
Require moderate temperatures to grow
Hardening
The process by which tolerable stress levels trigger the plant to prepare for extreme conditions
Changes in leaf when adapting to high temperatures
Surface: hairs can reduce evaporation and reflect intense sunlight
Size: smaller, thicker leaves help minimize effect of high light intensity
Angle/position: some species (legume) can moves leaves to reduce exposure to sunlight
Transpiration: increasing transpiration can help maintain the plant cooler than surrounding environment
Shading: plants use some structures to protect others from direct sunlight
Avoidance strategy: in hot environments, many species are short-lived annuals (ephemeral) avoiding worst conditions
Adaptations to cold temperatures
Rosette/low growing plants: protected against weight of snow and environment warmer close to ground
Evergreen plants: high concentrations of sugar in leaves acts as anti-freeze
Deciduous plants: leave abscission and bud scales to protect axillary meristems
Photosynthesis
A process that requires carbon dioxide and water and reactants and produces sugars and oxygen, requires sunlight
Chlorophyll
Captures sunlight for photosynthesis, molecules located in the thylakoid membranes in the chloroplasts
Light as an environmental stressor
Too low: not enough to cover plant’s metabolic requirements
Too high: unused energy leads to oxidative stress
Pulvini
Join-like thickening at base of leaves (legumes) that allows for movement of leaves (this is how plants change the position of their leaves to adjust to light intensity!!)
In ferns, mosses, and some seed plants chloroplasts move to different parts of the cell to adjust light intensity
Sun leaves
Smaller and thicker, with will-developed mesophyll and several layers of palisade cells, epidermal hairs might minimize excess sunlight
Shade leaves
Larger and more tender, with weakly developed mesophyll, epidermal cells might act as lenses to increase light capture
Mesophyll
Inner tissue of plant leaf, contains chloroplasts, primarily responsible for photosynthesis
Sun leaves at subcellular level
Chloroplast with small grant and few stromal thylakoids, photosystem: small proportion of antenna and many reaction centers
Shade leaves at sub cellular level
Chloroplast with large grant and many stromal thylakoids, photosystem: large proportion of antenna and few reaction centers
Stolons (runners) (spread)
Aboveground stems, emerge from crown and bend downwards, develop plantets where they touch the ground
Stoloniferous rhizomes (spread)
Belowground stems, upright shoots, develop from axillary buds in nodes, adventitious roots in underside
Suckers (spread)
Upright shoots growing from horizontal roots (aka “adventitious shoots)
Twiners (climb)
Stems of vines that grow in spiral around objects such as small trunks, supports itself
Tendrils (climb)
Modified leaf or short stem that wraps around objects they touch, provide support for main stem
Adhesive padsÂ
At tips of tendril-like short stems, allow plants to attach to walls
Supportive rootsÂ
In soft, wet soils plants form these to help them grow taller
Buttress roots (growing tall)
Wedge-shaped, tall and wide, prevent shallow-rooted trees from falling over
Prop roots (growing tall)
adventitious/aerial roots that arise from the trunk (or large branches) and grow downwards to provide support in shifty conditions (ex: mangroves)
Epiphytes (stealing light)
Grow in wet areas, use their roots to grasp tree branches (instead of water absorption)
Lianas
Typically in rainforests, woody, creeping vines that spread through the canopy spreading their leaves to the sun
Raunkiaer’s approach
Proposed a sequence of life forms depending on how they protect their meristem