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what do plants require? (4)
CO2, H2O, light energy and inorganic nutrients from environment
definition of essential nutrient for plant
it cannot complete its life cycle without them
micro and macro nutrients are classified based on...
the amount present in plant tissues
what are the three principal structural elements of plants?
C, H, and O
C, H, and O account for ___% of a plant's mass
96
What is phytoremediation?
plants help to clean toxic sites by uptake of heavy metals
what are the principal nutrient-absorbing structueres of a plant?
roots
assimilation in plants
apoplastic and symplastic pathways
apoplastic pathway
water and dissolved minerals diffues into the epidermal cells into the cortex
symplastic pathway
minerals cross root hair membrane and diffuse from cytoplasm to cytoplasm toward the stele
nitrogen fixation
Process of converting nitrogen gas (N2) into ammonia (NH3) or NH4+
- reduction
nitrification
the oxidation of N2 to NH3 or NH4+ to nitrite and nitrate
NH3 is
ammonia
NH4+ is
ammonium ion
NO2- is
nitrite
NO3- is
nitrate
what does nitrogen fixation?
bacteria and cyanobacteria
two enzymes that make up nitrogenase
dinitrogenase reductase and dinitrogenase
what determines if a plant can use a mineral (4)?
availability, binding vs leaching, assimilation, rhizosphere interactions
what is the industrial nitrogen fixation process called?
Haber-Bosch process
how does the Haber-Bosch process work?
N2 and H2 combined under high temp and pressure in the presence of a catalyst
the nitrogen fixation step is a oxidation/reduction rxn?
reduction
what do plant roots secrete to attract bacteria?
phenolic compounds
what is inhibitory to nitrogenase?
oxygenase
what is the importance of leghemoglobin?
it binds oxygen and controls oxygen levels within nodule cells and deliver O2 to bacterioids
most of phosphorus on earth is stored in
sediments and rock
plant strategies for improving phosphorus availability (3)
- root exudates (H2O, carboxylic acid, phosphatases) to enhance uptake
- cluster roots
- mycorrhizal associations
ectomyccorhizae grow how?
between root cells down to the endodermis
endomyccorhizae grow how?
between and inside cells down to the endodermis
what form must N2 be converted in order to be accessible to plants?
NH4+ or NO3-
what enzyme catalyzes N2 fixation?
nitrogenase
how much ATP does nitrogenase need to fix N2?
18 per nitrogen fixed
what is the major source for fertilizer?
mining of phosphate-containing rocks
what is translocation?
solute movement through phloem
does translocation move up or down?
down
translocation is a ______ stream
assimilate
what hypothesis explains translocation?
pressure flow hypothesis (Munch hypothesis)
what is transpiration?
water movement through xylem
does transpiration move up or down?
up
transpiration is a ______ stream
transpiration
what hypothesis explains transpiration?
cohesion-tension theory, cohesion-adhesion theory, transpirational pull
what is adhesion?
attraction between unlike molecules
what is cohesion?
attraction between like molecules due to hydrogen bonding
what are the three possible pathways for water movement through plants?
apoplastic, symplastic, and transcellular
apoplastic water movement
movement between cells and along cell walls
symplastic water movement
movement from cell to cell via plasmodesmata
transcellular water movement
movement through the membranes of cells, not necessarily through plasmodesmata
what happens if there is a break in the water column? cavitation?
perforation plates are set to trap embolisms, and pits which allow the by-pass of a blocked tracheary element
what forms the water channels in plants?
tracheary elements
what is an embolism?
air bubbles that form in the water column
what is the driving force of water transport in plants?
- cohesion of water molecules to eachother
-adhesion of water molecules to walls of vessel elements, resisting downward movement
- pull of evaporation from stomata
what kind of pressure is associated with translocation?
positive
what kind of pressure is associated with transpiration?
negative
when there is an increase in sugar content, water moves into the ______________ during translocation
sieve tube elements
turgor pressure is highest in the...
initial sieve tube element
what do phytohormones regulate?
cell division, elongation and differentiation
- also pattern formation, organogenesis, reproduction, sex determination and responses to stress
What does pleiotropic mean?
influence many traits
wide ranging effects
hormone steps
initial sensor, synthesis, transport, perception/receptor, signaling and responses
auxin
-promotes cell elongation, root formation and bud growth
- phototropism and gravitropism
what ways can hormones be made?
isoprenoid, amino acids, and lipids
the auxin hormone blocks the formation of what?
lateral branches
how do phytohormones move?
through the phloem or transporter proteins
auxin travels (transport)
down, from tip of shoot to tip of root
what is basipetal transport?
when auxin changes direction and moves short distances up the root again
what was the first experiment to discover auxin?
Darwin & Darwin
at low auxin levels, what happens? (signaling pathway)
Aux/IAA proteins and ARF protiens associate and interfere with ARF action, inhibiting transcription
the majority of phytohormones are
small organic molecules
what was the first phytohormone to be discovered?
auxin
at high levels of auxin...
Aux/IAA proteins associate with SCF complex, leading to proteolysis
- stops them from interfering with ARF protien
what does gibberellins do?
inhibits growth inhibitors
- seed germination, flowering, fruit growth, sex determination
gibberellins was first identified in...
fungal compounds
what do DELLA proteins do?
inhibit growth by blocking transcription
gibberellins triggers DELLA protein....
proteolysis
cytokinin functions
- cell multiplication and maintenance of shoot meristem
- also control of leaf senescence and nutrient allocation
how do auxin and CK act antagonistically?
auxin promotes root branching and CK promotes shoot branching
- CK promotes stem cell fate at SAM, auxin promotes lateral organ at SAM
- CK promotes differentiation at RAM, auxin maintains stem cell fate at RAM
ethylene promotes
senescence of leaves and petal
fruit ripening
abscisic acid
- stress responses
- control of stomatal aperture
what hormone accumulates in maturing seeds?
ABA
what hormone contributed to the evolution of drought tolerance in land plants?
ABA
seed germination requires the production/synthesis of what hormone?
GA
seed germination requires the catabolism/elimination of what what hormone?
ABA
what is the first downstream event from the binding of auxin or gibberellin to the respective receptors?
protein destruction
what hormones are known as inhibitors of an inhibitor?
auxin and GA
what hormones promote cell division and growth of the fruit?
auxin and GA