Chapter 2: A closer Look

FMIII Casket opening and closure methods

RAD141 CH 1 and 2 Clover Learnings

The Sonographer: A Closer Look

Clover Qs

biology unit 6(nitrogen cyle) new The Nitrogen Cycle Higher Tier Only Nitrogen is present as N2 gas in the atmosphere and within biological molecules, e.g. proteins, in the tissues of living organisms Nitrogen is cycled through ecosystems by the processes of the nitrogen cycle Uptake of nitrogen by living organisms N2 in the atmosphere is made available to living organisms by the process of nitrogen fixation Nitrogen fixation is carried out by nitrogen-fixing bacteria which convert N2 gas into ammonium compounds; these compounds are converted into nitrates in the soil Nitrogen-fixing bacteria can be free-living in the soil or they can live within root nodules of legume plants, e.g. peas, beans and clover Nitrogen gas can also be fixed by lightning when it strikes the earth, or during the production of chemical fertilisers After nitrogen fixation has occurred plants absorb the nitrates in the soil and use the nitrogen to build plant proteins Transfer of nitrogen between living organisms Animals feed on plants and digest the proteins in the plant tissues, providing nitrogen to build animal proteins Nitrogen may then be passed from one consumer to another up the food chain in the same way Release of nitrogen from living tissues Nitrogen from living organisms is returned to the soil in the form of ammonia by the action of decomposers such as bacteria and fungi When animals and plants die the proteins inside their tissues are broken down by the action of decomposers and returned to the soil in the form of ammonia Waste, i.e. urine and faeces, from animals contains urea, which is converted into ammonia by the action of the bacterial enzyme urease The plants can’t absorb ammonia so nitrifying bacteria convert the ammonia to nitrates which can then be taken up again by plants The conversion of ammonium compounds to nitrates is known as nitrification Returning nitrogen to the atmosphere Nitrates in the soil can be converted back into nitrogen gas (N2) by the action of denitrifying bacteria This process is known as denitrification Denitrifying bacteria are active in anaerobic conditions, e.g. in waterlogged or compacted soil Farmers can decrease the activity of denitrifying bacteria by ploughing the soil to increase aeration Nitrogen cycle diagram nitrogen-cycle-gcse The nitrogen cycle involves nitrogen fixation, decomposition, nitrification and denitrification Factors affecting the nitrogen cycle Because so many processes within the nitrogen cycle are carried out by microorganisms the cycle can be affected by factors that affect microorganism activity, e.g. Temperature This affects the rate at which enzyme-controlled reactions can occur Oxygen availability Aerobic bacteria rely on oxygen for respiration Low oxygen availability may lead to an increase in the activity of anaerobic bacteria, e.g. denitrifying bacteria pH This affects the rate of enzyme-controlled reactions as extreme pH levels can cause denaturation Water Water is needed by living organisms, so the rate of microbial activity increases in soil where moisture is present The presence of heavy metals in the soil Heavy metals, e.g. mercury and lead, can be toxic to the metabolism of microorganisms These factors are known to influence the rate at which decomposition occurs in compost heaps and landfill sites

Suture and Wound Closure Terminology

1 - Agonistes/Antagonistes des canaux chlorures

→ Understand what occurred at the Yalta Conference, including who was present at this meeting -“How do we deal with defeated or liberated countries of Eastern Europe after the war? -Pushed for interim government authority that represents a democracy -USSR would join the war against Japan when Germany is defeated → Claimed they would hold free elections in places freed from Nazi Germany, these free elections did not happen *Memebers present*: FDR (U.S), Churchill (U.K), Stalin (U.S.S.R) → Understand what was discussed at Potsdam, including who was present at this meeting and why that is important. Who is the consistent person at both conferences? → Post-War Germany→ What to do with it? -Confirmed plans to demilitarize Germany and split it into 4 occupied zones -Nuremberg trials tried Nazi’s for war crimes -Potsdam Declaration → Threatened massive attack on Japan unless they surrendered and laid out non-negotiable terms for peace → How did they split Germany? How did they split Berlin? -Split Germany into 4 occupation zones (France, U.S., U.K., U.S.S.R) → Split Berlin into 4 zones, giving the U.S.S.R the majority since Berlin lays in their zone of occupation → Berlin Wall splits Capitalism to the West and Communism to the East Capitalism vs. Communism (Week 10, 4/13: “Capitalism vs. Communism Packet.”) → Understand the fundamental differences between Capitalism and Communism → What major countries are Communist or Capitalist? Communism: USSR, China, North Korea, Vietnam Capitalist: Everyone else (U.S., France, G.B, South Korea) → What is the goal of both economic systems? → Communism: Eliminate social classes, create a classless/moneyless society → Private property is nonexistent, everything is public → Capitalism: Create a free market where individuals have the ability to make their own money and have independence to make their own economic decisions → Private property fuels the capitalist system Arms Race (Week 11, 4/20. “Arms Race PPT Reading”, “Arms Race PPT Dropbox”) → What is an “Arms Race?” Who participated in this “Race?” When two or more countries increase the size of military resources to gain military/political superiority over each other → U.S and U.S.S.R are the countries participating in this Arms Race → Start of the Cold War? → Dropping the nuclear bomb officially started the Arms Race → What type of Arms were being produced by major world powers? Nuclear weapons, ICBMs → How does an Arms Race affect the world/humanity in the world? MAD → If everyone is producing more powerful weapons this puts the world at risk for the possibility of Nuclear War. → If one launches a missile/drops a bomb then the other country retaliates until the world is basically over *The world is at a constant state of fear of the possibility of the unknown and potential end of the war* Space Race (Week 11, 4/22: “Space Race Primary Source Chart”) → What does the “Space Race” imply? Who is racing to space? -Who can get to Space first? The U.S or Soviet Union → Started with the race to space, developed into the race to the moon once the USSR reached space first… U.S reached the moon first → How did the Space Race affect U.S and Soviet Union relations? Greatly intensified Cold War tensions and the constant battle for superiority in any/every way possible for the US and USSR → Led to a greater technological race and a superiority aspect of it for both countries which further increased the need for national defense and funding for specific programs (NASA) Korean War (Week 11, 5/24: “Korean War Webquest”, Week 12, 4/27: “Korean War Textbook Analysis” ) → Who fought in the Korean War? Why was there a conflict in Korea? North Korea, the USSR, and China (communists) vs. South Korea and U.S → Korea used to be under Japanese influence until the end of WWII where it got split up at the 38th parallel by the US and USSR → The war started by the North invading the South and pushing them back to as far south as they could go before the U.S got involved and pushed them back to the 38th parallel and then some → Amercia feared that communism would continue to spread throughout the rest of the Asian continent → What is the legacy of the Korean War today? What is the dividing line between the two Koreas today? North Korea is still communist, South Korea is still capitalist → DMZ (demilitarized zone) at the 38th parallel that is still present today → Nobody won the war, both sides lost? → Forgotten war → Too many casualties → How did the Korean War symbolize the overall theme of the Cold War? This showed how the United States and Soviet Union were at a constant battle with one another and how there is the global struggle between communism and capitalism → Also demonstrated how concerned the United States was with the possible spread of Communism outside of the Soviet Union and how they would do anything to stop the spread Cuban Revolution (Week 12, 4/28: “Cuban Revolution Readings/Guided Questions.”) → Who was the Dictator of Cuba before Castro? Why did he have good relations with the United States? -Castro started to move away from U.S businesses and work more closely with the USSR who supported him. After the Cuban Revolution, the USSR was extremely supportive to Castro and the Cubans → When Castro came to power, how did Cuban relations with the U.S change? How did this change affect the Cuban relationship with the U.S.S.R? -Castro started to move away from U.S businesses and work more closely with the USSR who supported him. After the Cuban Revolution, the USSR was extremely supportive to Castro and the Cubans → Because the U.S openly disliked Castro and were not fans of him due to the change in American business AND the fact that Cuba was turning into a communist nation. → Castro nationalized land and businesses that were owned by the U.S which hurt their economy -Because of the strained relationship with the United States, this strained their relationship and strengthened the relationship between Cuba and the Soviet Union Bay of Pigs (Week 12, 5/1: “Bay of Pigs Packet”) → What was the “Bay of Pigs?” Who initiated this invasion? What was the overall goal of the United States by carrying out this plan? Was it successful? -Bay of Pigs was a failed attempt by the United States to overthrow Fidel Castro -They trained Cuban exiles who opposed Castro’s government to stage an uprising planned by the CIA → Goal was to overthrow Castro and put their own leader in control to benefit U.S business interest *Not successful* Invasion lasted 2 days and this pushed Cuba even closer to the USSR and made Castro look even stronger in Cuba Cuban Missile Crisis (Week 13, 5/4: “Cuban Missile Crisis Primary Source Worksheet”) → What is the “Cuban Missile Crisis?” How did tensions rise so high to almost reach a nuclear war? The U.S had missiles (ICBMs) in Italy and Turkey, close enough to fire at the USSR. The Soviet Union had missiles stationed in newly communist Cuba, 90 miles off the coast of Florida. The threat of a Nuclear War was as close as it ever was because nobody knew when a missile would launch, or who would be the one to launch it. → The U.S put a blockade on Cuba, which to the USSR seemed like an act of war. This increased the tensions between the two countries of who would “blink first” → Where did the U.S have missiles? Where did the U.S.S.R have them? U.S: Italy and Turkey USSR: Cuba Vietnam War (Week 13, 5/8. “Vietnam War Intro Notes”, Week 14, 5/11: “Vietnam War Stations Activity”) → Understand who was fighting on behalf of North and South AND the roles of both fighting parties supporting the North and South Vietnamese, respectively. The United States fought with South Vietnam, sending several million men to fight. The U.S was much more direct in the conflict, providing materials, strategy, men, and other equipment -The USSR did not send as many troops to fight with North Vietnam, but they did provide military aid (weapons), and military/logistical support → Main goal was to spread communism more in Asia → What was the significance of the Gulf of Tonkin AND the Gulf of Tonkin Resolution and how that affected United States involvement in the Vietnam War? The Gulf of Tonkin is in North Vietnam. U.S. destroyers got attacked, allegedly by the North Vietnamese which gave President Johnson the justification to enter the Vietnam War -Gulf of Tonkin Resolution was the response by President Johnson to enter the war without an official declaration of war approved by Congress → This led to the draft of 18-26 year old male citizens → Why was the Tet Offensive significant to the Vietnam War? How did this affect how people in the United States viewed the Vietnam War as a whole? North Vietnamese and Communist “Viet Cong” soldiers attack the South, giving heavy losses to them. Public support is going down significantly in the United States since this was the first televised war, showing the people at home how horrible the truths of war really are and how it is not being won by the U.S like the President is saying → Understand what a PROXY WAR is and the DOMINO THEORY and how it relates to the Cold War relations of the U.S and U.S.S.R. Proxy War is a conflict where a country supports other groups, often people not in the country, to fight a war against a common enemy without directly engaging into the conflict itself → These supported groups (Korea, Vietnam) can recieve military aid, funding, or training → Goal is to exert influence → EXAMPLES: Korea and Vietnam Domino Theory → The theory that if one country falls to communism then a surplus of others will fall to communism. This is a fear of the United States trying to prevent communism from spreading all over the world, putting their global dominance at risk

Project Management Audit and Closure Lecture Notes

Suture and Wound Closure Comprehensive Review

Storage Closets

Closer look 2 and skill 2

A closer look 2 and Skill 2

A Closer look 1

Swift Functions, Closures, and Data Structures: Key Concepts for Students

ENV 226: Essential Ecology Final Exam Study Guide — om single-species thinking to the dynamics of many interacting ecies. A community is more even when all species have similar abundances. Diversity: A combined measure of richness and evenness. More diverse = more likely to pull multiple different species out of a 'hat'. Shannon Diversity Index (H′): The most common diversity index. Higher H′ = more diverse (high richness AND high evenness). Formula: H′ = –Σ(pᵢ · ln pᵢ), where pᵢ is the proportion of individuals in species i. Worked example If a community has 4 species, each at 25% (p = 0.25), then H′ = –[4 × (0.25 × ln 0.25)] = 1.39. If one species dominates (e.g., 70/10/10/10), evenness drops and H′ falls even though richness is the same. Why diversity matters — ecosystem function & services Ecosystem function: Biological, geochemical, and physical processes that take place within an ecosystem (e.g., productivity, nutrient cycling, decomposition, pollination). Ecosystem services: The benefits humans derive from ecosystems. Four major categories: Provisioning: food, water, timber, fiber Regulating: climate regulation, flood control, water purification Cultural: recreation, spiritual, aesthetic, educational values Supporting: soil formation, nutrient cycling, primary production How diversity affects function — mechanism Complementary resource use (niche complementarity): Different species use slightly different resources (e.g., water at different soil depths, nutrients at different times). A diverse community captures more of the available resources than any single species could, raising total productivity. Diversity–stability theory Compensation: Species respond differently to environmental fluctuations. When one species declines, another can increase and 'compensate,' keeping overall ecosystem function steady. Insurance hypothesis: A diverse community is more likely to contain at least one species with traits that help the ecosystem cope with change. Diversity acts as ecological 'insurance' against disturbance. Rules of community assembly — what determines diversity at a site Three filters act in sequence on the regional species pool to determine which species actually end up in a local community: Term Definition Dispersal Who can physically get there. Controlled by distance from source populations and by dispersal ability. Connects to the 'mass effect' / rescue effect — regional diversity (gamma) can rescue local diversity (alpha). Environmental filtering What species can tolerate the abiotic conditions (climate, soil, water, salinity). Example: Ponderosa pine will not survive in the Sonoran Desert — environmental filtering excludes it. Biotic filtering What species can coexist given interactions with other species (competition, predation, facilitation). Strongest where abiotic conditions are benign, because more species can be there to interact. Intertidal zonation paradigm — how the filters stack In rocky intertidal communities, abiotic stress (desiccation, wave action) sets the UPPER limit of a species' distribution — an environmental filter. Competition and predation set the LOWER limit — biotic filters. Take-home: environmental filtering dominates in stressful zones; biotic filtering dominates in benign zones. What maintains diversity Intermediate Disturbance Hypothesis (IDH): Diversity is highest at intermediate frequencies or intensities of disturbance. Low disturbance lets competitive dominants exclude others; high disturbance eliminates all but the most disturbance-tolerant species. The middle keeps both groups in the community. Positive species interactions (facilitation): When one species makes conditions better for another (e.g., a nurse shrub providing shade and moisture for seedlings underneath). Facilitation tends to INCREASE biodiversity, especially in stressful environments. 1.2 Succession Primary succession: Colonization of a substrate that has NEVER supported life (e.g., bare bedrock, new volcanic rock, glacial retreat). Soil must be built from scratch, typically by pioneers like lichens and mosses. Secondary succession: Recovery after a disturbance that left soil and some biological legacy behind (e.g., a cleared field, most wildfires). Much faster than primary succession because soil and seed bank persist. Pioneer species: The first species to colonize a disturbed or bare area. Typically fast-growing, high-dispersal, stress-tolerant organisms that modify the site so later-successional species can establish. Quiz-style example The Woodbury Fire burned so intensely on the Tonto NF that only bedrock remained. Recolonization of this area is PRIMARY succession — there is no soil or seed bank left to start from. 1.3 Ecological Energetics Energy: The currency of ecosystems. Most ecological energy originates from the sun as electromagnetic radiation and is stored in tissues (biomass). Trophic level: Organisms that share the same function in the food chain and the same nutritional relationship to primary sources of energy. Level 1 = producers; 2 = primary consumers (herbivores); 3 = secondary consumers (carnivores); 4+ = tertiary / apex predators. Autotroph (primary producer): An organism that produces its own food from inorganic sources — typically plants, algae, and some bacteria via photosynthesis. Consumer (heterotroph): An organism that obtains energy by consuming other organisms. Primary consumers eat producers; secondary consumers eat primary consumers; etc. Production: The rate at which new biomass is created by organisms in an ecosystem (units of mass or energy per area per time). Net primary production (NPP): Gross primary production (total photosynthesis) MINUS the energy plants use for their own respiration. NPP is what is actually available to herbivores. Assimilation and production efficiency Energy is lost at every step of the grazing food chain. Two key efficiencies describe where energy goes: Term Definition Assimilation efficiency (Energy assimilated / energy consumed) × 100%. Assimilated = consumed – egested (waste). Herbivores ≈ 20–50% (tough plant material); carnivores ≈ 80% (similar tissue chemistry). Production efficiency (Energy in new biomass / energy assimilated) × 100%. Endotherms (birds, mammals) are LOW (~1–3%) because most energy is burned as heat; ectotherms (insects, reptiles, fish) are HIGH (~10–50%). Worked example (assimilation efficiency) Eats 400 J, excretes 200 J as waste, puts 50 J into growth. Assimilated = 400 – 200 = 200 J. Assimilation efficiency = 200 / 400 = 50%. The 10% rule Roughly 10% of the energy at one trophic level is transferred to the next. The rest is lost to respiration, heat, and waste. This is WHY food chains are short (usually 4–5 links): there simply isn't enough energy left to support another level. 1.4 Food Webs A food web is many, connected food chains — a map of who eats whom across an entire community. In simple diagrams, arrows point from prey to consumer. Complex diagrams use plus/minus signs to show the direction of effect, and dashed lines to show indirect effects. Top-down control: Higher trophic levels (predators) limit the abundance of lower levels. Removing a top predator releases herbivores, which suppress plants. Bottom-up control: Lower trophic levels (nutrients, producers) limit higher levels. Adding nutrients increases plants, which increases herbivores, which increases predators. Trophic cascade: Indirect effects of a predator propagate down the food web. Classic example: wolves reintroduced to Yellowstone → elk browsing decreases → riparian willow and aspen recover → beavers return → stream ecosystems recover. 2. Ecosystems Ecosystem: A community of organisms PLUS their shared environment. Includes biotic components (plants, herbivores, carnivores, detritivores) and abiotic components (climate, soils, nutrients). 2.1 Ecological building blocks Ecological building block: An atom that (1) makes up organisms and (2) is relatively abundant. Key building blocks: C, H, O, N, P (and sometimes S) — collectively CHONP. Not building blocks: Silicon, aluminum, arsenic, tungsten — they may be abundant in the crust or used by some organisms, but are not core structural elements of life. Potassium is important biologically but is NOT a core 'ecological building block' in this course's sense. 2.2 Liebig's Law of the Minimum Growth is dictated not by the total resources available, but by the SCARCEST resource. The 'limiting nutrient' sets the ceiling on production; adding more of a non-limiting nutrient has no effect until the limit is raised. Application — nutrient pollution A coastal system receives 10 g N, 200 g P, 50 g C, and 20 g O per year as pollutants, and you know the system is N-limited. By Liebig's Law, adding MORE nitrogen is what will most change structure and function — even though phosphorus is arriving in larger quantities, it is not the limiting nutrient. 2.3 Eutrophication Eutrophication is the enrichment of an aquatic system with nutrients (especially N and P) from fertilizer runoff, wastewater, or atmospheric deposition. Process: Excess N fuels algal blooms → algae die and sink → microbial decomposition consumes oxygen → a hypoxic 'dead zone' forms → fish and invertebrates die. Once N is drawn down, the system can become P-limited; phosphorus mined for fertilizer keeps the cycle going. The Gulf of Mexico hypoxic zone is the classic example. 2.4 Nutrient cycles (N, C, P) Term Definition Nitrogen cycle N₂ in atmosphere is biologically inert. Nitrogen-fixing bacteria (free-living and in legume root nodules) convert N₂ → ammonium (NH₄⁺). Nitrification converts NH₄⁺ → nitrite → nitrate (NO₃⁻), the form most plants take up. Denitrification returns N₂ to the atmosphere. Humans roughly DOUBLED global N fixation via the Haber-Bosch process → fertilizer → eutrophication. Phosphorus cycle Largely a SEDIMENTARY cycle — no gaseous phase. P weathers from rock → soil → plants → consumers → back to soil → eventually to ocean sediments. Slow turnover at global scales; a critical component of DNA/RNA, phospholipids, bones, and ATP. Carbon cycle See dedicated section below. C moves among atmospheric, terrestrial, oceanic, and fossil pools. Photosynthesis pulls CO₂ out; respiration and combustion return it. 2.5 Ecotones and cross-ecosystem flows Ecotone: A transition zone between two ecosystems, exhibiting gradients in environmental conditions and a related shift in the composition of plant and/or animal communities (e.g., forest–grassland edge, estuary). Two factors determine how a flow of material/energy from one ecosystem affects another: Relative size of the systems — when the amount of something varies across ecosystems, the LARGER system has a bigger impact on the small system (e.g., a stream flowing into a small pond vs. into the ocean). Quality of the resource — rich subsidies (like salmon carcasses bringing ocean nutrients to streams) matter more than dilute ones. 2.6 Ecological state change & resilience Key components of ecosystems: STRUCTURE (what organisms are there and how they interact), FUNCTION (processes of energy and nutrient movement), and REGIME (which of several possible stable states the system is in). Alternative stable states: An ecosystem can exist in two or more contrasting conditions under the same environmental conditions (e.g., clear lake vs. turbid lake; forest vs. shrubland). Ecological state change (regime shift): A large, persistent, often abrupt shift in the structure and function of an ecosystem, triggered by crossing a critical threshold. Threshold / tipping point: The level of a driver (stressor) at which a system flips to a new state. Hysteresis: Once a system flips, simply reversing the driver does NOT restore the original state — the return path is different from the 'forward' path. Slow vs. fast drivers: Slow drivers (e.g., gradual warming, soil nutrient accumulation) build up until a fast driver (e.g., fire, storm) tips the system across the threshold. Perturbation: Any event (abiotic or biotic) that disturbs the ecosystem. Perturbations that cause regime change can be abiotic (fire, flood, drought) or biotic (pest outbreak, invasion). Resilience: The capacity of a system to absorb disturbance, adapt to change, and recover from adversity while maintaining its essential functions, structure, and identity. The ball-and-cup diagram Picture a ball sitting in a valley (cup) on a hilly landscape. The ball is the current state of the ecosystem; the cup is the 'basin of attraction' for that state. A disturbance pushes the ball; stabilizing (negative) feedback loops pull it back. Strong disturbance or a shrinking cup (loss of resilience) can push the ball over a hill into a NEW cup — that's state change. Negative (stabilizing) feedback loop: A change triggers a response that DAMPENS the change, keeping the system near its current state. Deepens the cup. Positive (amplifying) feedback loop: A change triggers a response that AMPLIFIES the change, pushing the system further from its current state. Flattens the cup and makes state change more likely. Applying resilience to conservation & restoration Manage for resistance — remove stressors that push the ball (exclude high-intensity grazing, reduce pollution). Manage for resilience — rebuild the 'cup' by re-establishing key species, nutrient cycling, and stabilizing feedbacks (planting perennial grasses, restoring hydrology). Passive restoration works when the seed bank, soil, and key species are still intact; active restoration is needed when the system has already crossed the threshold. 3. Landscape Ecology and Biogeography 3.1 Landscape ecology Landscape ecology: The study of spatial patterns of ecosystems and their ecological consequences — explicitly considers the arrangement of habitats across space and how organisms and materials move through them. Spatial elements Term Definition Patch A relatively homogeneous area that differs from its surroundings (e.g., a forest stand in a grassland). Generally the highest-quality habitat. Matrix The background land-cover type that surrounds patches (e.g., desert in Saguaro NP, or agricultural land around forest fragments). Corridor A linear feature connecting patches — allows movement of organisms, genes, and energy. Examples: riparian strips, hedgerows, engineered wildlife crossings (Oracle Road, Tucson). Ecotone See above — the transition zone between landscape elements. Spatial heterogeneity Variability in environmental conditions and habitat types across a landscape. Drives diversity at landscape scales. Scale dependence Ecological patterns and processes depend on the spatial/temporal scale at which they are observed (e.g., a species may look stable regionally but be declining locally). Fragmentation Fragmentation breaks a large continuous habitat into smaller, more isolated patches. Effects include: Loss of total habitat area More edge relative to interior — edge effects (different microclimate, invasives, more predators) penetrate into remaining patches Reduced connectivity — animals cannot move between patches Smaller populations in each patch → inbreeding depression, loss of genetic variability, higher extinction risk Saguaro NP example Mid-sized carnivores in Saguaro NP West crashed after a disease outbreak and never recovered. Why? The city of Tucson grew between Saguaro NP East and West, severing connectivity. No recolonization could occur from the eastern population. Solution: re-establish connectivity — the Oracle Road wildlife crossings documented over 4,400 crossings by 16 species in their first two years. Patch dynamics Patch size, shape, and connectivity change over time because of ecological processes — succession, disturbance (fire, flood, windthrow), and fragmentation — not random chance and not just geology. 3.2 Biomes and realms Biome: A large biological community defined by climate and dominant vegetation type (e.g., tropical rainforest, boreal forest, tundra, desert, savanna, temperate grassland). Biogeographic realm: A large area of the Earth's surface with a distinctive assemblage of taxa, reflecting shared evolutionary history (e.g., Nearctic, Neotropical, Palearctic, Afrotropical, Indomalayan, Australasian, Oceanic, Antarctic). Factors shaping where biomes are found: temperature and precipitation (the primary controls), seasonality, latitude, elevation, continental geography, and evolutionary history. Realms reflect plate tectonics — Pangaea split into Laurasia and Gondwana, then into the continents we have today, producing unique evolutionary trajectories in each realm (e.g., Australia's marsupials, Madagascar's lemurs). 3.3 Island Biogeography and the SLOSS debate MacArthur & Wilson's Theory of Island Biogeography: species richness on an island is set by the balance between the colonization rate (immigration) and the extinction rate. Size effect — larger islands have LOWER extinction rates (bigger populations). Distance effect — islands closer to the mainland have HIGHER colonization rates. Equilibrium species number occurs where colonization and extinction curves INTERSECT. SLOSS debate — Single Large Or Several Small? Originally framed: is a single large reserve or several small reserves of equal total area better for biodiversity? Large favors: lower extinction, room for interior species, bigger populations, full food webs. Several small favors: replication (insurance against one disaster), sampling more habitat types, potentially higher total diversity. Modern answer: it depends — on species' dispersal, the matrix, and whether you value diversity vs. viability. Connectivity (corridors) often matters more than the large/small question alone. Source population: Produces more offspring than can be supported locally — exports individuals to other patches. Population growth rate > 0. Sink population: Organisms arrive but do not reproduce enough to sustain the local population; persists only via immigration from sources. Population growth rate < 0. 4. Extinction and Climate 4.1 The 'Big Five' mass extinctions Term Definition Ordovician–Silurian (~439 Mya) ~85% marine species lost. Cause: rapid glaciation and sea-level drop, then warming. Late Devonian (~364 Mya) Prolonged event; major loss of marine invertebrates, especially reef builders. Probable causes include ocean anoxia and climate change. Permian–Triassic (~251 Mya) 'The Great Dying' — ~96% marine species and ~70% terrestrial vertebrates. THE most severe. Cause: Siberian Traps volcanism → CO₂ spike → warming, ocean acidification, and anoxia. Recovery took 5–10 million years. End Triassic (~199–214 Mya) ~50% of species lost; cleared the way for dinosaurs to dominate. Likely cause: CAMP volcanism and climate change. Cretaceous–Tertiary (K-Pg, ~65 Mya) ~76% of species, including non-avian dinosaurs. Cause: Chicxulub asteroid impact (plus Deccan Traps volcanism) → darkened skies, cooling, then warming. Why scientists are concerned now Current extinction rates are 100–1000× background rates — comparable to mass-extinction levels. Rate of change: current climate change is occurring more rapidly than almost any past episode — faster than many species can adapt or track. Humans have built roads, cities, and agricultural landscapes that BLOCK the range shifts species would otherwise use to follow their climate. Human societies are themselves adapted to current climate (agriculture, supply chains, coastlines) — disruption drives conflict. 4.2 Why climate change affects ecological systems Temperature, precipitation, seasonality, and extreme events all drive the distribution and performance of every species. Shifting climate disrupts energy balance, water balance, food availability, and reproduction; changes the timing of seasonal events; and alters disturbance regimes (fire, floods, storms). All of these cascade through communities and ecosystems. 5. Climate Change — Ecology, Climate, and the Carbon Cycle 5.1 The carbon cycle Term Definition Pool (reservoir) A place where carbon is stored and from which it can be released. Measured as a quantity (e.g., gigatons). Flux The amount of carbon exchanged between pools per unit time (gigatons/year). Measures MOVEMENT. Sink A pool that accumulates more carbon than it releases — net REMOVER of carbon from the active cycle. Source A pool that releases more carbon than it accumulates — net ADDER of carbon to the active cycle. Biggest/smallest pools & fluxes Major carbon pools (approximate, gigatons): Deep ocean: ~37,000 GtC — BY FAR the largest pool Fossil pool (oil, gas, coal): ~10,000 GtC — second largest Reactive ocean sediments: ~6,000 GtC Soils: ~2,300 GtC Surface ocean: ~1,000 GtC Atmosphere: ~800 GtC — this is the pool that drives climate Plant biomass: ~550 GtC (the largest LIVING pool) Major fluxes are photosynthesis and respiration (~120 GtC/yr terrestrial; ~90 GtC/yr ocean), which are normally nearly balanced. Fossil-fuel combustion and deforestation are the (smaller but crucial) fluxes currently unbalancing the system. Why atmospheric CO₂ is increasing Humans are burning fossil fuels — moving carbon from a long-term sink (the fossil pool) into the active atmospheric pool faster than natural sinks can remove it. Deforestation and land-use change also shift carbon from plant biomass and soils to the atmosphere. The balanced photosynthesis/respiration fluxes cannot keep up with the ~10 GtC/yr added by human activity. 5.2 Ocean acidification As atmospheric CO₂ rises, more CO₂ dissolves into the ocean. The chemistry: Step 1: The ocean is slightly alkaline; CO₂ is slightly acidic, so CO₂ dissolves into seawater. Step 2: CO₂ + H₂O → H₂CO₃ (carbonic acid). Step 3: H₂CO₃ dissociates → HCO₃⁻ (bicarbonate) + H⁺. Step 4: Some HCO₃⁻ dissociates → CO₃²⁻ (carbonate) + H⁺. Step 5: Bicarbonate and carbonate exist in equilibrium. Net result: more H⁺ ions → lower pH = acidification. Acidification also reduces carbonate availability, making it harder for corals, shellfish, and plankton to build calcium-carbonate skeletons. Warming and the ocean's ability to sequester carbon Warmer water holds LESS dissolved CO₂ (inverse solubility). As oceans warm, their ability to absorb atmospheric CO₂ decreases — a positive feedback loop that further increases atmospheric CO₂ and warming. 5.3 Important climate feedback loops Term Definition Ice-albedo feedback (POSITIVE) Warming melts polar ice → darker ocean/land replaces reflective white ice → lower albedo, more solar energy absorbed → more warming → more melting. Water vapor feedback (POSITIVE) Warming increases evaporation; water vapor is a greenhouse gas → more warming → more evaporation. Permafrost/methane feedback (POSITIVE) Thawing permafrost releases CO₂ and CH₄ long locked in frozen soils → more warming → more thawing. CO₂ fertilization (NEGATIVE, partially) Higher CO₂ can boost plant photosynthesis, pulling more C out of the atmosphere. Partially counteracts warming but is limited by water, nutrients, and heat stress. Ocean solubility feedback (POSITIVE) Warmer oceans hold less CO₂ → more stays in the atmosphere → more warming. Quiz-style example Melting polar ice caps → decreased albedo → further warming = POSITIVE feedback loop (amplifies the original change). 5.4 Factors affecting Earth's surface temperature Three major controls: Energy arriving from the sun (solar radiation) Earth's albedo — how much of that energy is reflected back to space Greenhouse gases in the atmosphere — how much outgoing infrared is trapped Carbon dioxide is the LARGEST driver of current human-caused climate change (sheer volume, long atmospheric lifetime). Methane is more potent per molecule but far less abundant; water vapor amplifies change via feedback but is not itself a primary driver. 6. Climate Change — Ecological and Human Response 6.1 How climate change affects plants and animals Climate change disrupts performance in three main ways: Term Definition Energy balance Plants: respiration rates rise faster than photosynthesis with warming — net carbon gain (and growth) drops. Animals: thermoregulation costs rise; outside the thermal neutral zone, organisms burn more energy just to stay alive. Water balance Warmer temperatures and higher vapor-pressure deficit mean plants LOSE more water per unit of photosynthesis. Animals face greater dehydration risk; aquatic species face altered hydrology. Food acquisition & reproduction Changed phenology, drought, and heat reduce the resources available for growth and reproduction. Fewer seeds, fewer offspring, lower survival. Examples of species already affected Term Definition Pika Small alpine mammal restricted to cold, rocky talus. Warming pushes them to higher elevations — eventually they 'run out of mountain.' Already extirpated from lower-elevation sites in the Great Basin. Tuatara Reptile with temperature-dependent sex determination. Warming skews sex ratios toward males, threatening population persistence. Wolverine Depends on persistent spring snowpack for denning. Declining snowpack reduces suitable reproductive habitat. 6.2 Responses of species: MOVE, ADAPT, or DIE Move: shift range poleward or upslope to track suitable climate (classic response). Range shifts are highly variable across species — depends on dispersal ability, habitat specificity, and whether barriers (cities, roads, water bodies) intervene. Adapt: through plasticity (phenotypic change within a lifetime) or evolutionary change (genetic change across generations). Long-lived species with small populations adapt slowly. Die: local extirpation or global extinction if neither option is available fast enough. 6.3 Phenology Phenology: The timing of recurring biological events — bud burst, flowering, migration, breeding, hibernation. Climate change is advancing many spring phenological events (earlier bloom, earlier migration). Phenological mismatch occurs when interacting species shift their timing differently — e.g., a migratory bird arrives after its caterpillar prey has already peaked. Mismatches cascade through food webs. 6.4 Characteristics of climate-vulnerable species Narrow thermal tolerance (specialists) Poor dispersal ability (can't move to new climate) Long generation time, low reproductive rate (slow to adapt) Small, fragmented populations (low genetic variation, high stochastic risk) Dependence on climate-sensitive habitats (snowpack, sea ice, coral reefs, alpine tundra) Narrow geographic range, especially on islands or mountain tops (nowhere to go) Tightly tied to a specific phenological window or species interaction 6.5 Why current climate change is especially damaging Rate — change is occurring faster than most species can adapt or move Barriers — human land use has fragmented habitat, blocking the range shifts species used during past climate changes Cumulative stressors — climate change interacts with pollution, invasive species, overharvest, and habitat loss Interconnected systems — ecosystems, human agriculture, and global supply chains are all calibrated to current conditions 6.6 Mitigation vs. Adaptation Term Definition Climate MITIGATION Actions that reduce the magnitude of climate change itself — typically by lowering atmospheric greenhouse gases. Examples: switching to renewables, reforestation (sequestering carbon), reducing fossil-fuel use, more efficient buildings and transport. Climate ADAPTATION Actions that help humans and ecosystems COPE with the climate change that is already happening / unavoidable. Examples: creating migration corridors, building climate-resilient ecosystems through forest thinning, adjusting USDA seed zones, changing crop choices, updating hunting/fishing regulations, designing for sea-level rise. Quick quiz check Planting trees to sequester carbon = MITIGATION (reduces atmospheric CO₂). Thinning Southwest forests to make them more fire-resilient = ADAPTATION (copes with changing fire regime). Geoengineering proposals like stratospheric aerosol injection = a controversial form of mitigation (reduces incoming solar energy). Special cases of adaptation Managed (assisted) relocation: Actively moving species to areas outside their current range that are projected to become climatically suitable. Benefits: may be the only option for species that cannot disperse fast enough; can save species from extinction. Risks: recipient communities may experience novel interactions; potential to create invasive species; ethical questions about intervention. Assisted evolution: Human intervention to increase the rate of evolutionary adaptation — e.g., selective breeding for heat tolerance, or hybridization with warm-adapted populations. Benefits: keeps species in place; works for species that cannot move. Risks: may reduce genetic diversity; unintended consequences; can go wrong (outbreeding depression). 6.7 Corridors, climate refugia, and conservation design Climate refugium: A location whose physical or biological features allow species to persist despite regional climate change — e.g., high-elevation cool pockets, deep canyons, shaded slopes, coastal fog zones. Incorporating corridors (to enable range shifts) and refugia (places species can hold on) into reserve design is essential for climate-integrated conservation. A high-elevation forest that remains cool despite regional warming can serve as a seed source for recolonization — that's the textbook example of a refugium supporting resilience. Final thoughts: making an argument about climate-integrated conservation You should be able to give your own opinion on climate-integrated conservation and defend it. A solid answer acknowledges trade-offs: traditional 'protect what is there' approaches may fail under rapid change, but aggressive interventions (managed relocation, assisted evolution) carry real risks. Most conservation scientists argue for a portfolio approach — protect refugia, build corridors, and use active interventions only where the alternative is extinctionl

Closer look 2 and Skill 2

Closer look 1

Monnght - CLOSURES