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The Urogenital System is really two systems:
Urinary System and the Reproductive System.
Urinary System and the Reproductive System develop from
intermediate mesoderm
will help the body to maintain homeostasis by removing nitrogenous wastes, controlling blood volume, regulating blood pressure, maintaining the appropriate blood pH balance, and osmoregulating to control salt-fluid levels.
kidneys
The vertebrate kidney has a basic pattern of
of Glomeruli, Renal Tubules, and a pair of Excretory Ducts.
kidneys in all vertebrates will be located between the parietal peritoneum and the muscles roofing the coelom. This position is termed
Retroperitoneal
The glomerulus is responsible for
filtration aka first stage of urine formation
what is responsible for filtration aka first stage of urine formation
glomerulus
amniote glomerulus aka
renal corpsucle
renal corpsucle made up of
starting point of the renal tubule, the Bowman’s or Glomerular Capsule, and a rete mirabilia called the Glomerular Capillaries
capillaries of renal corpuscle use what for control of blood
afferent and efferent arteriole
a cup shaped structure that wraps around the glomerular capillaries.
glomerular capsule
here will be two layers the glomerular capsule separated by a space into which the filtrate enters. This space is called the
Urinary or Capsular Space.
The outer layer of the glomerular capsule is the
parietal layer
inner layer of glomerular capsule is visceral layer and is composed of unique epithelial cells termed
podocytes
The podocytes have primary and secondary processes that wrap around the glomerular capillaries. The secondary processes are the
pedicles
There will be small spaces between adjacent pedicles called
Filtration Slits
So the filtration membrane in the amniote glomerulus consists of
fenestra of the glomerular capillaries, the shared basement membrane, and the filtration slits.
an increase in blood pressure within the glomerular capillaries will force larger items out of the blood and across the filtration membrane. The product of this filtration is the
Glomerular Filtrate.
Glomerular Filtrate. This type of glomerulus is called an
Internal Glomerulus
. Earlier in development for most vertebrates there is another type of glomerulus, the
External glomerulus
e External Glomerulus. In this type of renal structure the glomerular capillaries are or are not in contact with the renal tubule.
are not
a funnel-shaped opening of the renal tubule called a
Nephrostome
Typically the nephrostome will have cilia that beat so as to establish a current drawing the glomerular filtrate into the renal tubule. These nephrostomes are termed
Ciliated Nephrostomes.
The renal tubules will be associated with a network of capillaries in most vertebrates, the
Peritubular Capillary Bed
The glomerular filtrate will have valuable materials (ex; glucose) that were filtered out of the blood recovered from it by crossing the walls of the renal tubule and entering into the surrounding capillaries. This process is called
Tubular Resorption
To facilitate the three processes of urine formation (filtration, tubular resorption, tubular excretion) the renal tubule and associated vasculature form a single functional unit,
Nephron
The kidneys arise from a portion of the splanchnic mesoderm termed the
Lateral Plate or Nephrogenic Mesoderm.
The hypothetical ancestral vertebrate kidney is termed the
Archinephros
e archinephric kidney is believed to have been several pairs of external glomeruli connecting to pairs renal tubules by way of the nephrostomes. These renal tubules connected to the two excretory ducts that carry the waster product to the cloaca termed the right and left
Archinephric Ducts.
n. The only thing that exists today that is close to the archinephric kidney is the
Holonephric Kidney
Holonephric Kidney develops into
Pronephric Kidney
Which type of glomerulus is suspended in the coelom and uses ciliated nephrostomes to draw filtrate into the renal tubule?
External glomerulus
What is the functional unit of the kidney that includes the glomerulus, renal tubule, and associated capillaries?
Nephron
Which structure draws filtrate from the coelom into the renal tubule in an external glomerulus?
Nephrostome
the first embryonic kidney in all other vertebrates
Pronephric Kidney
What are nephrostomes?
Funnel-like openings on pronephric tubules that draw filtrate from the body cavity.
Do pronephric kidneys last in the adult?
No, they are temporary.
What does the pronephros do while developing?
It induces the formation of the next kidney, the mesonephro
head kidney
pronephric
mesonephros is sometimes referred to as the
middle kidney because forms caudal to pronephros
The only remaining portion of the pronephros will be the pronephric duct which will at this point be called the
Mesonephric Duct
During the development of the mesonephric kidney the nephrogenic mesoderm will produce a new set of urinary tubules, the
Mesonephric Tubules
mesoderm is the part of the embryo’s mesoderm (middle layer of cells) that will develop into the kidneys.
nephrogenic
true or false: The mesonephros is the adult kidney in anamniotes but it a temporary structure in amniotes
true
do hagfish and lamprey display renal portal system
no
The mesonephric tubules in many jawed fishes and amphibians develop more caudally than in other anamniotes and may reach the area of the cloaca. For this reason the mesonephric kidney of these vertebrates is also known as the
Opisthonephric Kidney aka the rear facing kidney
in male jawed fish some of the more cranial mesonephric tubules are modified to carry
sperm
mesonephric tubules in male jawed fish are now termed the
vasa efferentia
explain what sexual kidney is
cranial mesonephric tubules in male jawed fish
The portion of the mesonephric duct that drains the sexual kidney in male jawed fish will be the
epididymis
s. To decrease the mixing of sperm and urine the mesonephric duct will be supplemented or replaced as an excretory organ by a pair of new ducts, the
accessory urine ducts
During the time period that the mesonephros is the functional kidney a new kidney will begin to form, the
metanephros
s. In females the mesonephric duct will degenerate into a series of blind ended tubules within the peritoneum associated with the ovaries. These are the
Eoophoron, Paroophoron, and Gartner’s Ducts
functional kidney of the adult amniote.
metanepheric
An extension of the mesonephric duct near the cloaca will extend into the nephrogenic mesoderm. This extension is termed the
metanephric bud
The stalk-like metanephric bud will become what of the kidney
ureter
ureter of kidney will expand to become
renal pelvis
The metanephric kidney will be covered by a fibrous connective tissue capsule called the
renal capsule
Due to its pattern of development the metanephros will have a depression on the medial aspect called the
hilus
what is the point of communication between kidney and ureter, renal vasculature, and nerves
hilus
internally kidney is divided into
renal cortex, renal medulla, and renal pelvis
The cortex will have extensions running deep into the medulla termed the
cortical or renal columns
Due to the renal columns the renal medulla will be divided into triangular segments called the
renal or medullary pyramids
These renal pyramids will contain portions of the nephrons, excluding the glomeruli, and a series of tubular ducts are a continuation of the collecting ducts called
papillary ducts
which pelvis is the innermost segment of the metanephros and is in direct contact with the ureter from which it arose.
renal
receives urine from the papillary ducts of the medulla and conveys it to the ureter.
renal pelvis
amniote kidney has the pelvis subdivided into the
Minor Calyces and the Major Calyces.
highly perforated area called thepapillary ducts in the renal pyramid will open at the apex giving rise to a
area cribrosa
the amniote ureters are paired organs that will convey to the
urinary bladder
The nephron begins in the cortex with the glomerulus. The glomerular filtrate will pass into the first segment of the renal tubule proper
Proximal Convoluted Tubule (PCT)
The proximal convoluted tubule will straighten and dive into the medulla to form the
proximal straight tubule PST
. The proximal straight tubule will connect with another straight segment called the
distal straight tubule
The distal straight tubule will ascend back to the cortex. The two straight segments form a loop called the
loop of henle
The distal straight tubule will drain into another twisted tubular structure called the
distal convoluted tube
The distal convoluted tubules will have a portion that sits against the glomerulus. This forms the
Juxtaglomerular Apparatus.
Birds and mammals have two classes of nephrons
Cortical Nephrons and Juxtamedullary Nephrons
juxtamedullary nephrons are longer, extend deeper into 544 the medulla, and have a second capillary bed in addition to the peritubular capillaries. This second bed is the
vasa recta
a terrestrial adaptation that aids in conserving water
urinary bladder
mucosa will be arranged into folds when the bladder is empty. These folds are the
rugae
In amphibians, reptiles, birds, and monotreme mammals the urinary bladder opens into the
cloaca
The kidney’s excretory ducts, mesonephric ducts in amphibians and ureters in amniotes, will also open directly into the
cloaca
species lack a cloaca, they will possess a new excretory organ, the
urethra
Unlike females, the urethra in males
both excretory and reproductive
bony fish urinary bladders are evaginations of
nephorgenic mesoderm instead of cloacal evagination
glandular organs that have both an exocrine and an endocrine role
gonads
gonads exocrine role is to produce
gametes
gonads produce which gametes in males
spermatazoa
gonads produce which gametes in females
ova
endocrine role of gonads is to
produce sex hormones
male gonad is
testis
female gonad
ovary
Both the male and female gonads develop from a pair of embryonic thickenings of the coelomic mesoderm located medial to the mesonephric duct. This structure is called the
genital/gonadal ridge
portion of genital ridge where primordial cells become established
germinal epithelium
As the developing gonads grow they become associated with a fold of the dorsal peritoneum. This peritoneal fold is the _________ in testicular development and the _______ in ovarian development
Mesorchium
Mesovarium
In testicular development what will form in the gonadal primordia
Primary Sex Cords
primary sex cords ivade deep into genital ridge and develop
seminiferous tubules
site of spermatogenesis
seminiferous tubules
in females what forms after primary sex cords
secondary sex cords
release of eggs
coelem into oviducts
release of sperm
lumen of the seminiferous tubules and enter into a series of ducts that will convey these gametes to the external environment