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Disturbance
Physical or environmental event that disrupts the structure or function of an ecosystem
Removes biomass
Disturbance initiates succession!
Without disturbance, communities shift to climax species!
Fires to floods, high winds, or landslides

Dispersal (and types of Dispersal)
The movement of individuals (or their propagules, such as seeds, spores, or larvae) away from their place of origin
Pioneer Species
Intermediate Species
Climax Community Species
Pioneer Species
Good at dispersal
Thrive in poor/absent soils
Help build soils
Short lifespan
Lots of offspring
Rapid growth
Not good competitors
Intermediate Species
Moderate dispersal
Require some soil nutrients
Help build soils more
Climax Community Species
Require rich soils
Great competitors
Slow growth
Good competitors
Long lifespan
Low dispersal rates
Not as many offspring
Not tolerance of harsh conditions
Succession and Stages
The process by which the community of species and habitat in an area changes over time
The sequence of plants and animals that appear at a site after a disturbance
Early Succession Habitat
Challenging physical conditions with high light and temperature, poor soil nutrients, few soil fungi, low soil moisture, but low competition
Mid-Succession Habitat
Intermediate state with both physical and biotic challenges available
Late-Succession Habitat
Challenging biotic conditions with intense competition for light and soil nutrients, but more tolerable physical conditions

Life-History
The sequence of life events involving growth and reproduction—essentially, how species allocate energy and other resources to survival and reproduction
Allocating resources is about growth OR reproduction OR defense OR maintenance
Every individual has a limited amount of time and energy to devote to the activities that are required to stay alive and reproduce → trade off!

(Fitness) Trade-Off
An inevitable compromise between traits that can't both be optimized
No species or strategy is best in all situations, and advantages always come with costs.
Traits that improve performance in one context often reduce performance in another.
Whether a strategy is successful depends on who else is present and what strategies they are using.
Tradeoffs can prevent any single species or strategy from dominating, allowing coexistence
Ecological systems shaped by tradeoffs often show dynamic outcomes, not fixed winners.
Chance plays a role

How do trade-offs allow organisms to allocate limited resources?
Energy budget
Energy must be allocated to different uses
Metabolism, Growth, Movement/dispersal, Reproduction, Defense
Investing more in one trait means investing less in another
