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how should sampling be carried out
in a manner which minimises the impact on wild species and habitats
techniques for sampling
point count, quadrats and transects (plants or slow moving organisms), capture techniques( nets and traps) for mobile species, camera traps and scat sampling (elusive species)
Hazards in fieldwork
Hazards in fieldwork include:
Control measures in field work
Control measures include appropriate:
How can identification of an organism in a sample be made?
classification guides, biological keys or analysis of DNA or protein
how can organisms be classified
taxonomy and phylogenetics
taxonomy
the identification and naming of organisms and their classification into groups based on shared characteristics. classic classification is based on morphology.
Phylogenetics
the study of evolutionary history and relationships among individuals or groups of organisms. it is changing the traditional classification of many organisms.
divergent evolution
when two or more species sharing a common ancestor become more different over time
convergent evolution
Process by which unrelated organisms independently evolve similarities when adapting to similar environments
how does phylogenetic work?
using heritable traits such as morphology, DNA sequences, and protein structure to make inferences about an organisms evolutionary history and create a phylogenetic tree
phylogenetic tree (phylogeny)
a diagrammatic hypothesis of its relationships to other organisms
taxonomic groups
Nematodes, arthropods and chordates
example of nematode model organism
c.elegans
example of arthropod model organism
drosophila melanogaster ( fruit fly )
example of chordates model organism
mice, rats, zebrafish
what gives information on environmental qualities such as pollution
presence, absence or abundance of indicator species
mark and recapture formula
N=MC/R
mark and recapture formula key
a sample of population is captured and marked (M) and released. after an interval of tie a second sample is captured (C). If some of the individuals are recaptured (R) then the total population (N) can be calculated
what does mark and recapture assume
all individuals have an equal chance of capture, that there is no immigration or emigration, and that individuals that are marked and released can mix fully and randomly with the total population.
methods of marking
Banding, tagging, surgical implantation, painting and hair clipping
measurements used to quantify animal behaviour
latency, frequency and duration
latency
the time between the stimulus occurring and respond behaviour
frequency
the number of times a behaviour occurs within observation period
duration
length of time each behaviour occurs during observation period
ethogram
lists species specific behaviours to be observed and recorded in the study.
anthropomorphism
the attribution of human traits, emotions or intentions to non - human entities. can lead to invalid conclusions
how to calculate time budget with ethogram
record duration of each of the behaviours in the ethogram, together with the total time of observation
evolution
is the change over time in the proportion of individuals in a population differing in one or more inherited traits.
during evolution changes in allele frequency occur through..
non random process of natural selection and sexual selection and random process of genetic drift
variation is the result of…
mutation
what is the original source of new sequences of DNA
mutation
what is selection pressures a result of
populations producing more offspring than the environment can support
result of individuals having variation that is better suited to environment
survive longer and produce more offspring which is passing on those alleles to next generation giving advantage
selection results in…
the non-random increase in the frequency of advantageous alleles and the non-random decrease in the frequency of deleterious alleles
sexual selection
the non random process involving the selection of alleles that increase the individuals chances of mating and producing offspring
sexual dimorphism
where two sexes of the same species exhibit different characteristics
sexual selection may lead to..
sexual dimorphism
sexual selection can be due to
female choice and male-male rivalry
female choice
involves females assessing the fitness of males. this may result in males attempting to 'attract' females through elaborate displays ( feathers, puffed chest)
male-male rivalry
large size or weaponry increases access to females through conflict ( antlers to compete for females)
how does genetic drift occur
when chance events cause unpredictable fluctuations in allele frequency's from one generation to the next
when does genetic drift have a larger impact
in smaller populations as alleles more likely to be lost from the gene pool.
genetics drift can occur because of..
bottleneck effect and founder effect
founder effect
occur through the isolation of a population from a larger population. the gene pool of the new population is not representative of that in the original gene pool.
bottleneck effect
occurs when a population size is reduced for at least one generation. the reduced population can have lower genetic diversity.
effect of genetic drift on new gene pool
it is altered because certain alleles may be under represented or over represented and allele frequencies change. ( new gene pool not representative or original one)
strong selection pressures =
rapid rate of evolution
selection pressures are
environmental factors that influence which individuals pass on their alleles
biotic selection pressures
competition, predation, disease, parasitism
abiotic selection pressures
changes in temperature, light, humidity, pH, salinity
hardy- Weinberg (HW) principle
in the absence of evolutionary influences, allele and genotype frequencies in a population will remain constant over the generations
conditions for maintaining the HW equilibrium
no natural selection, random mating, no mutation, large populations size, no gene flow through migration in or out
use of HW principle
determine whether a change in allele frequency is occurring in a population over time. changes suggests evolution is occurring
HW formula
p^2 + 2pq + q^2 = 1
HW key
p= frequency of dominant alleles (A)
q= frequency of recessive alleles (a)
p2=frequency of homozygous dominant genotype (AA)
2pq= frequency of heterozygous genotype (Aa)
q2= frequency of homozygous recessive genotype (aa)
fitness
an indication of an individuals ability to be successful at surviving and reproducing. it refers to the contribution made to the gene pool of the next generation by individual genotype
fitness can be defined in terms of
absolue or relative
absolute fitness
The ratio between the number of individuals of a particular genotype after selection, to those before selection ( after/before)
absolute fitness stable genotype
answer = 1.0
absolute fitness increase genotype
bigger than 1.0
absolute fitness decreased genotype
smaller than 1.0
relative fitness
the ratio of the number of surviving offspring per individual or a particular genotype to the umber of surfing offspring per individual of the most successful genotype
co evolution
the process by which two or more species evolve in response to selection pressures imposed by each other. a change in the traits of one species acts as a selection pressure on the other species seen with species that have symbiotic relationships
symbiosis
co evolved intimate relationships between members of two different species ( impact can be positive, negative or neutral)
types of symbiotic interactions
mutualism, commensalism and parasitism
mutualism
both organism are interdependent on each other for resources or other services, (+/+)
Commensalism
only one organism benefits (+/0)
parasitism
the parasite benefits in terms of energy or nutrients and the host is harmed in terms of loss of these resources (+/-)
the red queen hypothesis
in a co-evolutionary relationship, change in the traits of one species can act as a selection pressure on the other species, meaning that species in these relationships must adapt to avoid extinction.
sexual reproduction
the production of new living organisms by combining genetic information from two different individuals of different types (sexes)
costs of sexual reproduction
males being unable to produce offspring, only half of each parents genome passed onto offspring disrupting successful parent genomes
benefits sexual reproduction
genetic variation (benefits out weigh costs)
what does genetic variation provide
The raw material required for adaptation, giving sexually reproducing organisms a better chance of survival under changing selection pressures
red Queen hypothesis in terms of sexual reproduction
parasite pressure maintains sexual reproduction in the host population by selecting for the ability to produce rare genotypes that are resistant to infection
in co evolutionary relationships ( parasite and host) why are sexually producing hosts selected for
better able to resist and tolerate parasitism will have greater fitness,
benefit of host reproducing sexually
the genetic variability in their offspring reduces the chances that they will be susceptible to infection by parasites
asexual reproduction
a type of reproduction by which offspring arise from a single organism and inherit the genes of that parent only it does not involve the fusion of gametes
advantages asexual reproduction
entire genome passes on to offspring ( advantage in very narrow stable niches or when re-colonising disturbed habitats ) one parent can produce daughter cells and establish colony of unlimited size over time. offspring produced more often and in larger numbers
examples of asexual reproduction in eukaryotes
vegetative cloning in plants and parthenogenesis in lower plants and animals that lack fertilisation
parthenogenesis
reproduction from a female gamete without fertilisation. (more common in cooler climates which are disadvantageous to parasites or regions of low parasite density or diversity )
costs of asexual reproduction
not able to adapt easily to changes in environment, but mutations can occur that provide some degree of variation and enable some natural selection and evolution to occur
mechanisms that organisms that only use asexual reproduction have to increase variation
horizontal gene transfer
meiosis
the division of the nucleus that results in the formation of haploid gametes from a diploid gametocyte
homologous chromosomes
Chromosomes that have the same size, same centromere position and with the same sequence of genes at the same loci.
meiosis I
the chromosomes have replicated prior to meiosis 1, each consist of two genetically identical chromatids attached at the centromere, chromosomes condense and pair up. chiasmata form at points of contact between non sister chromatids of a homologous pair and sections of DNA are exchanged. (linked genes are those on the same chromosome crossing over can result in new combinations of alleles of these genes. ) the crossing over of DNA is random and produces genetically different recombinant chromosomes. spindle fibres attach to the homologous pairs and line them up at the equator of the spindle. the orientation of the pairs of homologous chromosomes at the equator is random. each pair of homologous chromosomes is positions independently of the pairs irrespective of their maternal and paternal origin. this is know as independent assortment . the chromosomes of each homologous pair are operated and move towards opposite pole. cytokinesis occurs and two daughter cells form .
meiosis II
each of the two cells produced in meiosis I undergo a further division which the sister chromatids of each chromosome are separated. a total of four haploid cells are produced.
chiasmata
the part of attachment of non sister chromatids of homologous chromosomes where crossing over takes place.