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what are environments
heterogeneous
how does variation occur
over space and time
what kind of scales does variation occur at
different spatial and temporal scales
what do organisms have
adaptations to deal with environmental variation
phenotype
traits of characteristics of an organism, influenced by genes and environment
fitness
ability of an organism to survive and reproduce
phenotypic plasticity
the ability of a single genotype to produce different phenotypes in different environments
example of phenotypic plasticity
gray tree frog tadpoles can sense predator chemical cues, they produce a phenotype that allows fast escape when predators are present and fast growth when predators are absent
what kinds of traits are plastic
behavior, growth, development, and reproduction
phenotypic tradeoffs
a phenotype experiences higher fitness in one environment and lower fitness in other environments
what can plasticity be beneficial in
variable environments
adaptations to enemies
many species alter their growth, body shape, and behavior in response to the presence of predators
example of adaptation to enemies
daphnia can produce helmets in the presence of predators
adaptations to enemies for plants
plants can response to herbivores via inducible defenses
example of plant adaptations to enemies
virgina pepperwood
adaptations to competition
plants can response to increased density by growing taller
example of plant adaptation to competition
jewelweed
variation in food availability
animals may alter behavior or morphology depending on food availability
example of variation in food availability
after a Burmese python eat it doubles the length of its intestine and increases heart size by 40% and blood flow by 10x
what does self fertilization allow
reproductive assurance but can result in inbreeding depression
what do some species exhibit
delayed selfing
what is an example of competition for mates
the hermaphroditic common pond snail delays egg laying if mates are unavailable, self fertilizing snails lay fewer eggs
adaptations to temperature variation
many organisms can adjust their physiology to maintain activity across different environmental temperatures
microhabitats
many animals respond to temperature by moving microhabitats
example of microhabitat
the desert iguana regulates its body temperature by basking on rocks, seeking shade, or burrowing in the ground
adaptations to water availability
in response to scarce water, many plants will close their stomata or grow longer roots
what do rocky tidal pools have
variable salinity (splashing waves, rain water)
what does copepod tigriopus synthesize
amino acids proline and alanine to balance osmotic potential with increased salt water
what is cost of adaptations to salinity variation
metabolic cost, increased respiration
adaptations to oxygen variation
less oxygen at high elevation
how do animals adjust to oxygen
animals can adjust breathing rate, heart rate, and number of red blood cells
migration
the seasonal movement of animals from one region to another
energy storage
animals accumulate fat or cache food supplied for periods of harsh weather when food is inaccessible
where do plants store nutrients and energy
in roots
dormancy
organisms dramatically reduce their metabolic processes
diapause (shutdown)
partial or complete physiological shutdown in response to unfavorable conditions; common in insects
hibernation (sleepy)
individuals reduce the energetic costs of being active by lowering heart rate and decreasing body temperatures; common in mammals
torpor (drowsy)
a brief period of dormancy in which individuals reduce activity and body temperature; common in birds and animals
aestivation (siesta)
shutting down metabolic processes during he summer in response to hot and dry conditions, examples include snails, desert tortoises, and crocodiles
what can cold response occur at
different intensities
compared to species living in a homogenous environment, a species living in a heterogeneous environment is more likely to
exhibit phenotypic plasticity
an organism that can surviving well in cold temperatures but not in hot temperatures illustrates the concept of
a phenotypic trade off
if self fertilization results in low fitness, why would an organism choose self fertilization as a mating strategy
it ensures reproduction in low population densities
which behavioral is optimal for an organism using central place foraging
increase time spent searching for food at longer distances