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step 1 of perceptual process
distal stimulus
distal stimulus
objects in the environment available to observer that they attend to, the stimulus impinges on receptors for an internal representation
step 2 of perceptual process
light is reflected and focused
proximal stimulus
when the eye focuses on the light and creates a 2D representation of the tree unto the retina
Principle of Transformation
When the stimuli and responses created by the stimuli are transformed, or changed, between the environmental stimuli and perception
camera obscura effect
when bright light passes through a small aperture into a small space, projecting an upside down copy.
step 3 of perceptual process
Receptor processes
receptor processes
rod and cone receptors line the back of the eye, change light energy into electrical energy and influence what we perceive. this results in electrical representation of the tree within the neural circuits of the retina
sensory receptors
cells specialized to respond to env energy
transduction
key event which changes env energy to nerve impulses, occurs during step 3 (receptor processes)
step 4 of perceptual process
neural processing
neural processing
the different processes that occur as signals are transmitted through the maze-like network of neurons from the retina to the brain
cortex primary receiving areas
occipital - vision
temporal - hearing
parietal - touch
step 5-7 of perceptual process
behavioral responses
behavioral responses
electrical signals transformed into conscious experience <→ person perceives object (perception)<→ recognizes it as a tree (placing it into a category)(recognition) <→ lets have a closer look (action)
knowledge
information the perceiver brings to a situation
bottom up/databased processing
processing based on incoming stimuli from the env
top down/knowledge based processing
processing based on perceivers previous knowledge
the intersection between topdown and bottom up processing
perception
psychophysics
showing someone something and then observing their behavior as we lacked sufficient physiological measures to peer into the brain
stimulus perception and stimulus physiology relationship experiment
neurophysiological activity should predict/correlate with perceptual performance; the finest line width that can be perceived by the subject indicates the subjects grating acuity
stimulus physiology relationship
copla - measures relationship between bar orientation stimuli and brain activity
oblique effect
phenomenon where oblique (angled) grating produces smaller levels of brain activity than vertical/horizontal grating
absolute threshold
smallest amount of energy needed to detect a stimulus
cross-over point
the threshold, found using method of limits (stimuli of different intensities presenting in ascending and descending orders which observer responds to whether or not they perceived the stimulus)
what is the identity of the stimulus
recognition testing
how quickly can i react to it
reaction time
How can i describe what is out there
phenomenological report
difference between physical and perceptual measures
subjects estimate that the magnitude is lower than it actually is
fovea rod and cone distributions
solely consists of cones
peripheral retina rod and cone distribution
both rods and cones, but more rods than cones
macular degeneration
fovea and small surrounding area destroyed, creating blind spot in center of field, common in older individuals
retinitis pigmentosa
genetic though everyone has it, rods destroyed in periphery then fovea attacked as well. in severe cases it leads to complete blindness
how many rods and cones are there in the eye
120 million rods to 6 million cones
the blind spot
where the optic nerve leaves the eye, on edge of visual field but is filled in from the other eye. located 12-15 degrees eccentricity, is 5.5×7.5 deg,1.5 deg below horizontal
cornea
fixed and unchanging; 80% of focusing
lens
adjusts shape for object distance; 20% of focusing when ciliary muscles tighten the lens thickens. light pass through lens more sharply, focus on objects near retina
near point
the closest focal point the lens can accommodate
presbyopia
the distance of the near point increases due to hardening of the lens and weakening of ciliary muscles → corrective lens needed for close activities
myopia
nearsightedness - inability to see distant objects clearly, focus point in front of retina. caused by refractive myopia: cornea or lens bends too much light or axial myopia: eyeball is too long
hyperopia
farsightedness - inability to see nearby objects clearly. focus point behind retina, usually caused by short eyeball. constant accommodation for nearby objects can lead to eyestrain and headaches
receptor cells
specialized cells in retina that convert light to electrical signals for visual perception
electromagnetic spectrum
a continuum of all possible forms of electromagnetic energy
what is most perceived light
reflected light, we are only able to see a very small range of existing wavelengths (380-740 nm)
phototransduction
the process of transferring energy from photons to the atoms and molecules of receptor photo-pigments in the retina, converting light into electrical signals for the brain to interpret as visual images
what happens during phototransduction
the retinal absorbs a photon (leads to isomerization)
Isomerization
the retinal changes shape after absorbing a photon, triggers the opsin to initiate an enzymatic chemical chain reaction (an enzyme cascade). isomerizing one pigment leads to the activation of a rod receptor
cascade reaction
single reaction leads to increasing numbers of chemical reactions
dark adaptation
the process of increasing sensitivity in the dark (measured by determining a dark adaptation curve)
process needed for transduction
retinal molecule changes shape
opsin molecule separates
retina shows visual pigment bleaching
retinal and opsin recombine to respond to light
visual pigment regenerates
light threshold
lowest in the middle of the spectrum, meaning most sensitive at around 580 nm
spectral sensitivity curve
1/threshold = sensitivity
rod spectral sensitivity (scotopic)
sensitive to short-wavelength light (500 nm)
cone spectral sensitivity (photopic)
3 types:
sensitive to short wavelength (419)
sensitive to medium wavelength (531)
sensitive to long wavelength (559 nm)
purkinje shift
enhanced sensitivity to short wavelengths during dark adaptation when the shift from cone to rod vision occurs
key components of neurons
cell body - dendrites - axon or nerve fiber
sensory receptors
specialized neurons that respond to specific kinds of energy
electrical signals in neurons
light energy enters the eye → (through the optic nerve) lateral geniculate nucleus in thalamus → visual receiving area (V1 or striate cortex)
recording electrical signals in neurons
2 small electrodes used to record from single neurons (1 inside and outside) difference in charge of -70 mV (resting potential)
resting potential
negative charge of neuron relative to its surroundings (-70 mV)
action potentials
show propogated response, 1 ms refractory prd (500-800 max upper firing rate), increase in stimulus intensity means increase in rate as size of action potential remains the same regardless of intensity. spontaneous activity of action potential occurs w/o stimulation
chem basis of action potentials
sodium flows into axon making it more positive (40 mV), positively charged potassium flows out of the axon making, charge returns to resting level
synapse
small space between neurons
neurotransmitters
released by presynaptic neurons from vesicles, received by postsynaptic neuron on receptor sites, matched into receptor sites, and used as triggers for voltage change in the post synaptic neuron
depolarization
caused by excitatory transmitters. neuron becomes more positive increasing likelihood of action potential
hyperpolarization
caused by inhibitor transmitters, neurons become more negative and decreases the likelihood of action potentials
convergence
process of multiple neurons sending signals to a single neuron to combine and process info from diff sources
high convergence
126 mill rods and cones to 1 mill ganglion cells (rods converge more than cones 120+6!!)
moderate convergence
120 rods and 6 cones to each ganglion cell
no convergence
1 cone to 1 ganglion cell (happens in fovea)
how does light pass through the eye
ganglion cells → bipolar cells → receptors
rod sensitivity
rods have greater convergence and sensitivity but cannot distinguish as much detail
Cone sensitivity
high visual acuity, incredible detail but need more light to respond than rods
lateral inhibition
neural mechanism in which an excited neuron reduces the activity of its neighbors, enhancing the contrast in sensory input
receptive field
the area of receptors that affects the firing rate of a given neuron in the neural circuit. determined by monitoring single cell responses
center on vs off cells
Con - light in center leads to highest firing rate
Coff - light in surround leads to highest firing rate
signal travel through the brain
lateral geniculate nucleus
primary visual receiving area in occipital lobe (striate cortex/V1)
2 pathways to temporal and parietal lobes
frontal lobe
major function of LGN
regulate neural information from retina to visual cortex, signals received from retina, cortex, brain stem, and thalamus. signals organized by eye, receptor type, type of env info
what cells are feature detectors
simple cortical cells (orientation)
complex cortical cells (orientation and direction)
end stopped cortical cell (orientation direction and length)
simple cortical cell
excitatory and inhibitory areas arranged side by side. responds best to bars of particular orientation
orientation tuning curve
shows how impulse is measured as a function of orientation for simple cortical cells
optic nerve fiber (ganglion cell)
center-surround receptive field responds best to small spots but will respond to other stimuli
lateral geniculate cell
center-surround receptive field similar to ganglion cell
complex cortical cell
responds best to movement of a correctly oriented bar across the receptive field. responds best to particular direction of movement
end-stopped cortical cell
responds to corners, angles, or bars of particular length moving in a particular direction
selective adaptation
neurons tuned to specific stimuli fatigue when exposure is long; firing rate decreases and fire less when stimulus immediately presented again. only neurons that respond to specific stimulus adapt
gratings
alternating light and dark bars
selective rearing
neural plasticity shows that when not shown some types of stimuli those types of neurons do not grow as much and thus lose ability to see these stimuli
prosopagnosia
inability to recognize faces, occurs due to damage in FFA (fusiform face area)
sensory coding
a representation of perceived objects through changes in neural firing of multiple individual neurons
specificity coding
specific neurons responding to specific stimuli, leads to grandmother cell hypothesis (each person gets own neuron)
problems - too many different stimuli to assign specific neurons
most neurons respond to a number of different stimuli
population coding
the pattern of firing across many neurons encodes specific objects, a large number of stimuli can be coded by a few neurons
sparse coding
only a small subset of neurons is active at any given time to represent specific stimuli
contextual modulation
response to stimulation within the receptive field can be affected by outside the receptive field
Topology
the study of geometric properties and spatial relations unaffected by the continuous change of a shape or the size of figures. refers to the way in which constituent parts are interrelated or arranged. topological maps are more concerned with relative than absolute positions
cochlea
tonotopic resonance along linear surface, representation of ears membrane, transduces sound and wired with receptors that have distinct topologu
auditory cortex
tonotopic map of cochlea surface
retinotopic map
electron map of the retina on the cortex
cortical magnification
small area of fovea is represented by large area on visual cortex, most of the visual field periphery has little computing power
positron emission tomography
radioactive tracer that moves through bloodstream and measures radioactivity in blood flow. Changes in blood flow → changes in brain activity