single-celled eukaryotes, either free-living or parasitic ex:amoeba
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porifera
sponges (think pores “pore” infera)
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Why are insects so diverse?
Small body size - There are many opportunities for small organisms in a given environment
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Why are insects so small?
1. It takes time to grow large - A long juvenile period increases the risk of mortality before reproduction 2. Oxygen limitation - Gas exchange becomes difficult at large sizes because of distance effects on oxygen diffusion
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Tympana
A thin cuticular membrane that allows for hearing in insects also mating calls
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Cnidaria
jellyfish, coral etc.
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sound
Pressure fluctuation transmitted in a wave-form via movement of air or water
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Hypothese on origins that have 4 overlapping stages
1. appearance of organic molecules 2. polymerization of molecules into RNA, DNA, or proteins 3. formation of protobionts 4. evolution of cell-like characteristics
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minimal properties of life
reproduction, metabolism, evolution
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hypotheses about how/where organic molecules originated
Absorption of nutrients, waste excretion, energy acquisition and transformation, and cellular synthesis
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Miller-Urey 1952 Experiment
Set up anaerobic atmospheric conditions and added an electrical discharge, resulting in the spontaneous formation of simple organic molecules
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Extraterrestrial hypothesis
Meteorites brought organic carbon to Earth
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Protobiont
Aggregate of molecules and macromolecules that acquired a boundary, allowing it to maintain an internal environment separated from its surroundings
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What makes protobionts possible precursors to living cells?
1. A boundary separated external environment from internal contents 2. Polymers inside the protobiont contained information 3. Polymers inside the protobiont had enzymatic function 4. Protobionts capable of self-replication
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living cells may have evolved from
coacervates-droplets that form spontaneously from charged polymers where enzymes trapped inside perform metabolic functions
liposomes-can enclose RNA
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what is the first macromolecule of protobionts
RNA
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what are functions of RNA
1. ability to store info 2. self-replication 3. enzymatic function (ribozymes)
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Radioisotope dating
A common way to estimate the age of a fossil by analyzing the decay of radioisotopes within the rock (usually igneous rock)
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Half-life
Length of time required for exactly one-half of original isotope to decay
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Factors that affect the completeness of the fossil record:
1. anatomy 2. size 3. number 4. environment 5. time 6. geological processes 7. paleontology
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fossil prokaryote
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cyanobacteria
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stromatolites
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changes in living organisms due to
genetic changes and changes in environment which allow for new organisms and extinctions
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Cretaceous
\- 144-65 MYA \n -Dinosaurs still dominant \n -Angiosperms - earliest flowering plants \n -Mass extinction at end wiped out dinosaurs and many other species
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Macroevolution
The formation of new species or groups of related species
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Lamarck
Proposed theory that organisms pass on traits they acquire in their lifetime. (Incorrect) i.e. Scars
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Charles Darwin
English natural scientist who formulated a theory of evolution by natural selection
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Darwin's Theory of Evolution based on:
1. Variation within a given species 2. Natural selection
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Population genetics
The study of genes and genotypes in a population
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Purpose of Population Genetics:
1. To know extent of genetic variation 2. Why it exists 3. How it is maintained 4. How it changes over the course of many generations
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Phenotype
Physical appearance or observed traits due to genotype
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Natural selection
A process in which individuals that have certain inherited traits tend to survive and reproduce at higher rates than other individuals because of those traits.
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Reproductive success
Likelihood of an individual contributing fertile offspring to the next generation
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Horizontal gene transfer
Transfer of genes between cells of the same generation
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Permian-Triassic Extinction Event
252 MYA, the only known mass extinction of insects \n - 8 or 9 orders became extinct
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Paleodictyopterida
An extinct superorder of beaked insects \n - Included 50% of all known Paleozoic insects
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Ultrasonic pulses
Method by which echolocating bats detect prey
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Phantom Echo hypothesis
Bat perceives clicks as multiple objects surrounding the moth
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Ranging Interference hypothesis
Bat perceives clicks as a blurred target
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Masking hypothesis
Bat perceives clicks as no target
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Four periods of the Earth
Archaen, Proterozoic, Phanerozoic, Cenozoic
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Life
An organized genetic unit capable of metabolism, reproduction, and evolution
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Reproduction
Growth, duplication of all cellular components, division of discrete cells
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Evolution
Variation, reproduction, and mutation
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Origin of Life: Stage 1
Nucleotides and amino acids spontaneously produced prior to the existence of cells (Primordial soup)
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Origin of Life: Stage 2
Nucleotides and amino acids became polymerized to form DNA, RNA, and proteins
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Origin of Life: Stage 3
Polymers became enclosed in membranes, separating them from the environment
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Origin of Life: Stage 4
Polymers enclosed in membranes acquired cellular properties
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Reducing Atmosphere hypothesis
Atmospheric reactions with lightning, solar, and cosmic radiation
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Deep-sea Vent hypothesis
Biologically important molecules may have been formed in the temperature gradient between extremely hot vent water and cold ocean water
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Coacervates
Droplets that form spontaneously from the association of charge polymers
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Liposomes
Vesicles surrounded by a lipid layer
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Key functions of RNA:
1. Ability to store information 2. Capacity for self-replication 3. Enzymatic function
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Sedimentary rock
Comprised of many layers. Sediments pile up and become rock
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Stromatolites
Fossilized biofilms, form in layers. Can be used to extrapolate fossilized prokaryotes
\-3.8 to 2.5 BYA \n -First cells are prokaryotic and heterotrophic \n -Little free oxygen, organisms were anaerobic \n -Autotrophs evolved as supply of organic molecules dwindled
\- 543-490 MYA \n -Warm and wet with no ice at poles \n -Cambrian Explosion - abrupt increase in diversity of animal species (unknown cause) \n -All existing major types of marine invertebrates that no longer exist
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Ordovician
\- 490-443 MYA \n -Diverse group of marine invertebrates including trilobites and brachiopods \n -Primitive land plants and arthropods first invade land \n -Towards end, abrupt climate change results in mass extinction (60% of marine invertebrates go extinct)
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Silurian
\- 443-417 MYA \n -Coral reefs appeared \n -Large colonization by terrestrial plants and animals \n -Spiders and centipedes \n -Earliest vascular species
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Devonian
\- 417-354 MYA \n -Ferns, horsetails, and seed plants (gymnosperms) \n -Insects emerge \n -Amphibians (tetrapods) emerge \n -Near end, series of extinctions eliminate many marine species \n -"Age of fish"
* 290-248 MYA \-Pangaea * Amphibians prevalent, reptiles dominant \-First mammal-like reptiles appeared \-Largest known mass extinction event at the end (90-95% of marine species)
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Triassic
\- 248-206 MYA \n -Reptiles plentiful \n -First dinosaurs and first true mammals \n -Gymosperms dominant land plant \n -Volcanic eruptions led to global warming and mass extinctions near end
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Jurassic
\- 206-144 MYA \n -Gymnosperms still dominant \n -Dinosaurs dominant land animals \n -First known bird \n -Mammals present but not prevalent
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Tertiary
\- 65-1.8 MYA \n -Mammals that survived expanded rapidly \n -Angiosperms became dominant land plant \n -Whales appeared \n -Hominoids appeared 7 MYA
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Quarternary
\- 1.8 MYA to present \n -Periodic ice ages cover Europe and North America \n -Certain hominoids become more human-like \n -Homo sapiens appear 130,000 years ago
When an exon (parts of a gene that encode protein domains) and the flanking introns are inserted into a gene
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Genomic level changes
Large changes in chromosome structure and number
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Allele
A variant form of a gene
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Genotype
Genetic makeup or hereditary information composed of multiple alleles
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Allele frequency
\# of copies of an allele/total # of alleles in a population
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Genotype frequency
\# of Indies with a particular genotype/ total # of indivs in a population
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Hardy-Weinberg equation
p2 + 2pq + q2 = 1
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Hardy-Weinberg conditions:
1. No new mutations 2. No natural selection 3. Large population - allele frequency doesn't change due to random sampling error 4. No migration between different populations 5. Random mating
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How do changes happen?
1. Introduction of new genetic information (mutations) 2. Mechanisms that alter the prevalence of alleles/genotypes