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Evolution
Change overtime in the genetic composition of a population
Aristotle classification
Species are fixed, arranged them on a scala naturae (ex. a latter with humans at the top and insects at the bottom)
The old testament classification
Species were individually designed by god
Carolus Linnaeus classification
Organismal adaptation; creator had designed each species for a specific purpose
Founder of taxonomy
branch of biology that classifies organisms
Carolus Linnaeus
Founder of taxonomy; binomial nomenclature
Levels of biological classification
Domain Kingdom Phylum Class Order Family Genus Species
Types of Domains
Eubacteria, Archaea, Eukarya
Georges Cuvier
Developed paleontology-study of fossils
Catastrophism
Each boundary between strata represents a catastrophe
Gradualism
Geologic change results from slow & gradual, continuous process
Uniformitarianism
Earth’s processes same rate in past & present (therefore Earth is very old)
Jean-Baptiste Landmark
Published Theory of Evolution (1809)
Inheritance of acquired characteristics
Modifications can be passed on
Thomas Malthus
Believed that consequences of overproducing within environment= war, famine, disease (limits of human pop.)
Charles Darwin
An English naturalist who was well known for his theory of evolution through natural selection
Alfred Russel Wallace
Also an english naturalist who independently covered the theory of evolution through natural selection
Darwinian View
History of life is like a tree with branches that represents life’s unity & diversity
Natural selection
Nature decides winners vs. losers
Artificial selection
Human modification of other species by selective breeding
Adaptations
Enhances an organisms ability to survive and reproduce
Fossil record
Provides evidence of the extinction of species, the origin of new groups, and changes within groups overtime
Homology
Similarity resulting from common ancestry
Homologous structures
Anatomical resemblances that represent variations of a structural theme
Vestigal structures
Remnants of features that served important functions in the organisms ancestors (ex.appendix)
Molecular homogies
Genes shared among organisms inherited from a common ancestor
Convergent evolution
The evolution of similar, or analogous features in distantly related groups
Analogous traits
Arise when groups independently adapt to similar environments in similar ways
Pangea
supercontinent that used to exist but split apart due to continental drift
Speciation
The process where one species splits into two species
Microevolution
Changes within a single gene pool
Macroevolution
Evolutionary change above the species level
Biological species concept
Defined by Ernst Mayr, population whose members can interbreed & produce viable, fertile, offspring; reproductively compatible
Distinct species
Songs & behaviors are different enough to prevent interbreeding
reproductive isolation
barriers that prevent members of 2 species from producing viable fertile hybrids
Prezygotic barriers
A reproductive barrier that prevents mating or fertilization (if mating occurs)
Geographical isolation
species occur in different areas; physical barrier
Ecological isolation
Species occur in same region, but occupy different habitats so they rarely encounter each other
Temporal isolation
species that breed during different times of day, different seasons, or different years
Behavioral isolation
Unique behavioral patterns & rituals isolate species
Mechanical isolation
morphological differences can prevent sucessful mating
Gametic isolation
Sperm of one species may not be able to fertilize eggs of another species
Biochemcial barrier
Sperm cannot penetrate egg
Receptor recognition
lock & key between egg & sperm
Chemical incompatibility
Sperm cannot survive in female reproductive tract
Postzygotic barriers
prevent hybrid zyogte from developing into viable adult
reduced hybrid viability
genes of different parent species may interact and impair the hybrids development
Reduced hybrid fertility
Even if hybrids are “vigorous”, they may be sterile
Hybrid breakdown
Hybrids may be fertile & viable in first generation, but when they mate offspring are feeble or sterile
The biological species concept cannot be applied to:
fossils & asexual organisms (including prokaryotes)
Allopatric speciation
Geographically isolated populations
Sympatric Speciation
Overlapping populations; reproductive isolation
Autopolyploid
Extra sets of chromosomes
Adaptive radiation
Many new species arise from a single common ancestor
Hybrid zones
Incomplete reproductive barriers
Reinforcement
strengthening of reproductive barriers
Fusion
Weakening of reproductive barriers
Discrete characters
Can be classified on an either or basis
Point mutations
Changes in one base
Chromosomal mutations
delete, duplicate, disrupt, rearrange; usually harmful
Sexual recombination
contributes to most of genetic variation in a population
Population genetics
Study of how populations change genetically overtime
Population
Group of individuals that live in the same area and interbreed, producing fertile offspring
Gene pool
All of the alleles for all genes in all the members of the population
Fixed allele
All members of a population only have 1 allele for a particular trait
Hardy Weinberg Principle
The allele and genotype frequencies of a population will remain constant from generation to generation UNLESS they are acted upon by forces other than Mendelian segregation and recombination of alleles
Equilibrium
Allele and genotype frequencies remain cosntant
Hardy Weinberg Principle equlibrium
p+q=1
p
frequency of dominant allele (A)
q
frequency of recessive allele (a)
Hardy Weinberg Principle equation
p²+2pq+q²=1
p²
AA (homozygous dominant)
q²
aa (homozygous recessive)
2pq
Aa (Heterozygous)
Minor causes of evolution
Mutations and Nonrandom mating
Major causes of evolution
Natural selection, genetic drift, gene flow
Natural selection
Individuals with variations better suited to environment pass more alleles to next generation
Genetic Drift
Small populations have greater chance of fluctuations in allele frequencies from one generation to another
Founder effect
A few individuals isolated from larger population; certain alleles under/over represented
Bottleneck effect
Sudden change in environment drastically reduces population size
Gene flow
Movement of fertile individuals between populations; gain/lose alleles; reduce genetic differences between populations
How does natural selection cause evolution?
Natural selection can alter the frequencies certain traits in a population in 3 ways
directional selection
phenotype of one extreme in a distribution is selected for
disruptive (diversifying) selection
Both extremes selected for
stabilizing selection
Middle phenotype is selected for
Sexual dimorphism
difference between 2 sexes (ex. size, color)
Intrasexual
selection within same sex (ex. males compete w other males for females)
Intersexual
mate choice (ex. females choose a colorful male)
Diploidy
hide recessive alleles that are less favorable
Heterozygote advantage
greater fitness than homozygotes
How old is Earth?
4.6 billion years old
When did the first life forms appear?
3.8 billion years ago
RNA World
The theory that the first genetic material was RNA
Sedimentary rock fossils
Layers called strata
Mineralized fossils
hard body structures
Organic fossils
rare in fossils but found in amber and frozen tar pits
Incomplete record fossils
many organisms not preserved, fossils destroyed, or not yet found
Relative dating
Uses order of rock strata to determine relative age of fossils