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What is heterotrophy
Organisms that depend on preexisting organic molecules for carbon and energy
What are fungi
Complex eukaryotic organisms that depend on organic molecules for energy, but are unable to digest food like animals
What are hyphae
Highly branched multicellular filament that provides a large surface area in fungi for nutrient absorption
What is mycelium
A network of branching hyphae
What is turgor pressure
A fungal material transport method involving molecules being taken up by the organism, driving water into the organism via osmosis, increasing pressure which is then decreased by growth and respiration due to molecule consumption
What is chitin
A modified polysaccharide containing nitrogen that makes up the cell walls of fungi and the hard exoskeletons of arthropods
What is a septum
A wall in fungal hyphae that partially divides the cytoplasm into separate cells
What are yeasts
Single celled fungi found in moist nutrient rich environments, that can metabolize sugar via fementation
What are ectomycorrhizae
One of the two main types of mycorrhizae; which produce a thick sheath of fungal cells (hyphae) that surround a root tip, and the hyphae that grow between cells in the interior of a root
What are endomycorrhizae
A mycorrhizae which penetrate into root cells where they produce highly branched structures (arbuscules) that provide a large surface area for nutrient exchange
What are endophytes
Fungi that live within leaves and may help the host plant by producing chemicals that deter pathogens and herbivores
What are lichen
Stable associations between a fungus and a photosynthetic organism usually being a green alga or a Cyanobacteria
What are fruiting bodies
Multicellular structures in some fungi that facilitate the dispersal of sexually produced spores
What is plasmogamy
The cytoplasmic unity of two cells
What is karyogamy
The fusion of two nuclei following plasmogamy
What does heterokaryotic mean
A stage in the life cycle of some fungi, in which plasmogamy is not followed immediately by karyogamy and the cells have infused haploid nuclei from both parents
What does dikaryotic (n + n) mean
Heterokaryotic cells that have two haploid nuclei, one from each parent in each cell
What is a mating type
A genetically distinct form of individual fungi in a species, that enables fertilization only between different types, to prevent self fertilization and promote gene crossing
What does parasexual mean
Asexual species that create genetic diversity via the crossing over of DNA during mitosis
What are chytrids
Aquatic fungi that have simple, unicellular bodies that produce gametes and spores with flagella
What are zygomycetes
A group of fungi that produce hyphae undivided by septa and do not form multicellular fruiting bodies
What are glomeromycetes
A group of fungi of low diversity but tremendous ecological importance that occurs in mycorrhizae
What are dikarya
A Fungal group that includes 98% of fungal species in which dikaryotic cells are formed
What are ascomycetes
A group of dikaryotic fungi in which nuclear fusion and meiosis take place in an elongated sac called an ascus (sac fungi)
What are basidiomycetes
A group of dikaryotic fungi in which nuclear fusion and meiosis take place in a club shaped cell called a basidium
What roles do fungi play in the environment
They decompose plant and animal tissues, play a critical role in the carbon cycle, and have ecological relationships with other organisms
Why are fungi necessary for the carbon cycle
They can locate and break down complex molecules and bulky tissue in other organisms
How do fungi have important ecological relationships
Some help increase plant growth on roots, some are agricultural pests (hergotism on wheat) , and some such as yeast are used to ferment plant carbohydrates to produce alcohol
How do fungi obtain energy
They are heterotrophs, but they can’t break down food like animals, so they absorb organic molecules directly (simple molecules pass easily, but complex molecules require enzymes to be broken down)
Why are hyphae important
The numerous thin hyphae provide fungi with a large surface area for nutrient absorption, allows it to find nutrients (due to no movement), and their rapid growth allows them to find new food patches as they grow
What is an interesting trait about mycelium
It can grow to become a large branching network, the largest covering over 2000 acres, and it can also grow to form fruiting bodies
What is turgor pressure similar to
Pholem transport in plants, due to pressure differences driving transport (concentration gradients)
What do fungi’s abilities to transport materials allow them to do
The hyphae can use materials in nutrient rich areas to fuel hyphae elongation across nutrient poor locations
What do septa do
They have pores that allow water and solutes to move between cells as well as the ability to plug them in case of injury
How do yeast reproduce
Asexually, by budding off
Where do most fungi receive nutrients
Dead organic material, which broken down contributes to the carbon cycle and places nutrients into the soil
How do fungi break down cellulose
Some fungi produce an enzyme to break down lignin (a fiber found in cellulose), which allows for easier decomposition
What kinds of fungal pathogens infect plants
Rusts, smuts, and molds infect plants
How are plant infections spread above ground
, by fungal spores carried by the wind or insects
how are plant infections spread below ground
Hyphae penetrate roots of plants
What are other ways plants get infected
Fungi infect plants through wounds, stomata, or by penetrating epidermal cells directly by degrading the wall with enzymes, then pushing hyphae into interior
How do fungal pathogens deal with invertebrates
They create hyphal rings that trap small organisms like C. Elegans
How do fungal pathogens deal with vertebrates
Many ways, such as toenail fungus, which are hard to treat due to them being eukaryotic
What are mycorrhizal relationships
Relationships between fungi and plants where the fungi supply plant roots with nutrients such as phosphorus and in return receive carbohydrates
Are mycorrhizal relationships between fungi and animals possible
Rare but yes, insects can provide fungi with shelter food, and protection in exchange for a food source
How do lichen reproduce
As a single organism, despite being an association between fungi and photosynthetic microorganisms
How are lichen built
Mostly of fungal hyphae, with an algae or Cyanobacteria forming a thin layer under the surface
How do hyphae help lichen
They help anchor lichen to a rock or tree, aid in water and nutrient uptake, and produce chemicals for defense against light and herbivores
How do microorganisms benefit lichen
They provide a reduced carbon source like a carbohydrate, as well as Cyanobacteria providing a source of nitrogen
Why are both organisms in lichen nessecary
They both exchange nutrients through hyphae that encircle or penetrate the cell walls, which allows lichen to survive as one in areas where neither could survive by themselves
What are the challenges of reproduction in fungi
Maintaining genetic variation and dispersal
How do fungi disperse in their life cycles
They produce spores that are carried by animals, the wind, or water (the aquatic spores have flagella)
What are properties of spore production
There is a low probability for proper spore location, so fungi produce huge numbers, these can be viable for long periods of time. They also allow fungi to use resources patchy in time and space, with this shortage cueing further production
How are spores formed asexually
They are produced with sporangia that are produced at the ends of hyphae, releasing spores into the air. Allowing the fungi to proliferate and disperse to new environments
What is used to spread sexual spores
Fruiting bodies on the surface, that are formed from densely packed hyphae at an elevation, leading to better dispersal
How do the shape of spores help
The football shape reduces drag and allows them to move fast
How does the asexual fungal life cycle work
Spores are produced by fungi, which are then dispersed and germinated, restarting the cycle
How do fungi sexually reproduce
Their hyphal tips grow together, and release enzymes that digest cell walls, allowing the hyphal cells to merge into a single cell with two nuclei
What is the sexual reproduction life cycle in fungi
Hyphal cells merge by the plasma, becoming dikaryotic, where later, the nuclei merge to form a diploid cell, which goes through meiosis to form spores that are dispersed to form new fungi
How do fungi accomplish sex without gender
Mating types, where fertilization can only take place with individuals with different alleles at the mating type gene
How are mating types increasing diversity
Fungal species have two or more mating type alleles, allowing for an increase in individuals they are able to mate with, increasing diversity
How does a parasexual cycle work
Nuclei in a heterokaryotic cell undergo karyogamy to form a diploid nucleus, where crossing over occurs in mitosis, leading to new gene combinations
How diverse are fungi
There are 75k known species not spread evenly through phylogeny, with 98% being dikaryotic
What are traits of chytrids
They are decomposers or pathogens found in moist environments, with single cells made of chitin that form short multinucleated structures, but lack well defined hyphae and a heterokaryotic stage, but form flagellated gametes that swim through their environment
What is the disease caused by chytrids
Chytridiomycosis caused by batrachochytrium dendrobatidis, killing amphibians
What are traits of zygomycetes
A small group of fungus, with some being decomposers (leaves, feces, food) , and others living on other organisms that grow mycelium and produce Ariel spores (Rhizopus, a black bread mold that digests simple carbon compounds)
How do zygomycetes reproduce
After karyogamy and meiosis, haploid cells produce an elevated stalk that spreads spores
What are traits of glomeromycets
They are monophyletic with low diversity, found with plant roots, increasing nutrient uptake for them
How do glomerocytes reproduce
Parasexual resulting in genetic diversity
What are traits of dikarya
Every mitotic division forms a new septum, allowing them to control the number of nuclei in each cell and proliferate
Includes edible mushrooms, yeast for beer bread and cheese, wood rotting fungi, and pathogens
What are differences in dikaryotic fungi
They evolved similar traits independently, as well as ascomycetes having a brief dikaryotic state compared to basidiomycetes having a longer state
How do ascomycetes reproduce differently
They release spores from a disc like ascus
How do ascomycetes affect human health
Ergot is a common pathogen, that when consumed can be used to treat migraines, but in high doses causes delirium and hallucinations
How can fungi influence behavior of animals
Some fungi can infect ants causing them to be beneficial to the fungus
What are traits of basidiomycetes
They make up 34% of fungi, and are made up of smuts, rusts, and a group that forms fruiting bodies
What makes the basidiomycetes life cycle different
It’s club shaped fruiting body is made up of only dikaryotic hyphae, and the products of meiosis do not undergo mitosis
How do gill shaped mushrooms spread spores
They drop from each gill from the basidum and are then moved into the atmosphere
What is coenocytic organization
An organism where the nucleus divides multiple times, but the cytoplasm does not , resulting in a large cell
What is cell adhesion
Molecules that cause adjacent cells to stick together, with little communication or transfer of resources between cells and little differentiation of specialized cell types (cells must stick together)
What is cell communication
Cells being able to communicate with one another
What is genetic programs
Cells participating in a network of genetic interactions for cell division and differentiation
What is bulk flow
Molecules moving at rates beyond those possible by diffusion across a concentration gradient
What is diffusion
Random motion of molecules with net movement from areas of higher to lower concentration
What is surface area to volume ratio
A necessary trait for cells to effectively transport molecules, with a high ratio making it easier
What are choanoflagellates
Single celled eukaryotes closely related to animals due to them having cell adhesion and cell signaling (contains genes that code for Cadherin and integrin proteins)
What is a meristem
Specific regions of growth in plants, found at the tips of roots and stems (due to cell wall and growth by division, expansion, and differentiation)
What is a blastula
A hollow, fluid filled ball of cells that develops from the morula and is an early stage of embryo development in animals
What is a gastrula
A stage of early embryo development in animals when cells of the hollow blastula migrate and reorganize to form a multilayered structure in which different cell types begin to differentiate
How many groups of eukaryotes are multicellular
36 out of 119
What are the other 83 groups of eukaryotes
Single celled organisms that eat other microorganisms or small organic particles, live suspended in water columns, or are parasites living within other organisms
Where do simple multicellular organisms originate
From colonial organisms, which stuck together for reproduction and feeding
What are traits of simple multicellular organisms
They contain adhesion molecules to allow cells to stick together, despite little communication, resource transfer, or differentiation, each cell has a full range of functions, and every cell is in contact with the environment for diffusion
What is an advantage of being multicelllular
Avoiding predation as predators cannot focus on one cell
What occurs in organisms with coenocytic organization
They are one large cell with multiple nuclei ass the cytoplasm does not divide, sometimes visible to the naked eye
What are the cons of multicellularity
Higher energy costs to develop, grow, and maintain tissues
Different functions require cells to cooperate (due to specialization during development using protein gradients)
Opportunities for individual cells to use resources to grow themselves and not the rest of the organism (cancer)
What does complex multicellularity require
Cell adhesion, cell communication, and genetic programs
What is three-dimensional organization
Multicellular organisms that have some cells in direct contact with the environment, meaning they need mechanisms for transferring environmental signals received by surface cells to interior cells, where genes will be activated in response
How many times did complex multicellularity evolve
6 times, once in animals, in green algae (leading to vascular plants), red algae, brown algae, and fungi twice
What is a key challenge of complex multicellularity
Transporting food, oxygen, and molecular signals rapidly across large distances within the body and exporting waste products out of the body
How are small distance limits via diffusion overcame by animals
A sponge places active cells close to the environment by using pores and canals
Jellyfish have thin layers of tissue, but have a thick shape due to inactive molecules
Animals have a high ratio of surface area to volume ratio in some cells, and use bulk transport