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kill me ( first 4 pp)
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Learning
a change in behavior as a result of experience
Learning is associated with ________ changes and the ______ capability to learn
neurological, genetic
Learning allows animals to
adapt to their environment
Learning results in
increased fitness
Should learning evolve in a fixed or dynamic world?
neither
Fixed world
no need to learn because of environmental constancy
Dynamic world
learning is not beneficial because of the unpredictable changes in the environment
The real world is ___
intermediate
Is learning adaptive?
Yes, it is and its expected to evolve
Evolution of learning is based on:
regularity of the environment, and reliability of past experience
Whats the simpliest form of learning?
habituation
Habituation
reduction and then lack of response to a stimulus over time.
Stimulus
can be anything in the environment
reaction
is a response
Nuerology
the synapse is believed to play an important role in learning and memory
the synapse
the junctions between two nuerons
IMHV
intermediate and medial parts of hyper striatum ventral
In birds IMHV plays an important role in
memory related to imprinting
memory
retention of a learned experience
Neural Plasticity
the structural changes in the brain. especially in the number and chemical strength of synapses between neurons
Synaptic connections chance due to
formation and elimination of dendritic spines
The dynamic nature of spines
may play a roll in learning
Stimulus-response association
making an association between an environmental stimulus and a behavioral response.
Classical Conditioning (Pavlovian conditioning)
Learning new associations between stimulus and an innate or unlearned response
absent of choice
Operant conditioning
Involves learning associations between learned behavior sand outcomes
Operant conditioning can measure
an animals progress toward solving a problem, the learning curve, a decline in errors over time
4 types of Operant conditioning
Positive reinforcement, negative reinforcement, positive punishment, negative punishment
positive reinforcement
a behavior becomes more likely due to the presentation of a stimulus, such as food.
negative reinforcement
a behavior becomes more likely due to the removal of a stimulus, such as pain
positive punishment
a behavior becomes less likely due to the presentation of a stimulus
negative punishment
a behavior becomes less likely due to the removal of stimulus.
the ability to learn has both
costs and benefits
costs can include
size/weight of brain, ability to specialize in other areas etc
Learning follows the rules of natural selections-
learning only evolves where learning is needed.
social learning develops when it
reduces the time and energy costs of learning alone
Social interactions
facilitates learning
Social learning with prairies dogs: How do young p dogs learn their predators and how to avoid them?
they learn antipredator behavior from adults
What faciliates this learning
local enhancement, public info
Local enhancement
an individual uses the presence of another as a cue for learning in the environment
public information
info obtained from activity or performance of others about the quality of the an environmental parameter or resource.
Social info in sticklebacks: high numbers of fish should indicate bets food patch
local enhancement
Social info in sticklebacks: feeding success of individuals should indicate best food patch regardless of number in the patch
public info
behavioral traditions
differences in behavior among pops, transmitted across generations through social learning ( culture)
Ex of behavioral traditions
song dialect, characteristic song difference that vary geographically among populations. common in birds
cognition
acquisition, storage, retrieval, and use of info
Cognition includes:
tool use, mirror self recognition, numerical competency, memory of past info, behavioral flexibility
Social Brain hypothesis
social pops evolve higher cognitive ability because of their social structure
Individuals that live in groups:
must recognize others and understand their social relationship to successfully interact with them in different ways.
Social brain hypothesi states:
that sociality can promote the evolutions of larger brains
foraging
acquiring food
four basic steps of foraging
Locate food
Select food
capture food
Consume food
Active foraging
involves search image and can use special mechanisms like heat and electricity
Passive foraging
(Sit-and-wait)
Elaborate capture mechanism, also specialized mechanisms
Passive foraging
mimicry- visual lores,heat sensors, mimicry- olfactory lures
time/energy
costs/benefits
Optimal foraging theory
Optimal behavior is the behavior that maximizes fitness based on natural selection.
The diet model assumes:
Foragers maximize fitness by. maximizing energy intake rate, food items are encountered on at a time in proportion to their abundance, food items can be ranked by their profitability.
Prey profitability
(j/s)=e1/s1+h1
E=
energy content
S=
Search/wait time
h=
handling
Diet Breadth
is affected by the abundance of the most profitable prey items, and not by the abundance of the least profitable items
Optimal diet model solution
is to always consume types A, B, and C while always ignoring types D and E
Zero-one rule
each food item is either always eaten or always rejected ( doesn’t actually help)
foraging decisions=
not always optimal
Optimal foraging questions
Where to look? Risk? How long to stay? Competition? When to eat something else?
Game of diminishing returns:
when a forager enters.a food patch, it initially harvest food at a high rate. But as the patch is depleted, its harvest rate declines
Optimal patch -use model
foragers experience diminishing returns as exploits a food patch.
the marginal benefit of feeding in a food patch is the instantaneous harvest rate.
Because of diminishing returns, the marginal benefit of feeding declines with time spent in a patch
The optimal patch-use model assumes:
Foragers attempt to maximize energy intake rate.
All patches are identical, travel time between patches are constant, The instantaneous harvest rate declines as a forager depletes a patch.
Optimal patch time is affected by
travel time
The optimal patch- use model predicts the
optimal time to spend exploiting each patch
for short travel times, the optimal patch time is
small
for longer travel times, the optimal patch time is
longer
Resource of defense
any feature that increases the amount of time spent by a predator in attaining is a
Sensory modalities
physical, chemical, morphological, and behavioral methods
Morphological
camouflage
physical defense
defenses which physically deter a consumer from obtaining a resource
Chemical defense
Plants produce a wide variety of toxic compounds, animals can make their own toxins but many consume toxic plants , then store the toxins themselves
Constitutive chemicals
are toxic only in high quantities and are always present
Inducible chemicals
are toxic in the smallest quantities but are only produced when attacked,
Morphological defenses
shape or color of an organism deters consumption
Cryptic morphology acts as
camouflage , either in color or body shape
Aposematic coloration
alerts consumers that an organism is toxic
Mimicry
takes on the colors/patterns of a toxic organism
Batesian:
The mimic itself is not toxic, only bluffing
Mullerian
The original and the mimic are both toxic , compounding their defense
Behavioral defense
Organisms modify their natural behaviors to avoid being consumed
Behavioral defense example
tunnels and burrow systems for movement, nocturnal active time, Herding or social groups, playing dead or false attack
Being social is a
behavioral defense
Active social defense include
mobbing, alarm calling, and vigilance behavioral
Passice social defense revolve around the
cost/benefit of gathering in large groups
confusion effect hypothesis
whole lot of animals, if they start running around then it s hard to pick
Dilution effect hypothesis
odds of surviving are better. 1/10 eaten stil 1/100 eaten
Selfish Herd hypothesis
you don’t have to be the fastest just faster than the slowest
Beneficial adaptation
when heritable variation and differential reproduction are present in a population, the trait that confers higher fitness spreads.
this trait is called beneficial adaptation becuase
it confers higher genetic success to individuals that possess it as compared to individuals that do not have it
Perfection isnt always adaptable
the right mutation doesn’t always arise
A single gene can have multiple
developmental consequences
Pleiotrophy
describes the fact that most genes have multiple developmental effcects
A gene might code for a
particularly useful adaptation against a predator, but also from a harmful developmental consequence