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Primary Succession
occurs on newly exposed substrates
Secondary Succession
occurs after disturbances
Facilitation mdel
early successional species prepare the way for late stage species facilitating their success
Inhibitation
early successionaal species hold the site against all invaders. THey make the site less suitable for other species. New species take over as past species die.
Tolerance model
Later stage species can invade and mature independent of species that precede or follow them. succession is associated with autogenic changes in environmental conditions
autogenic
environmental change resulting from presence and activities of organisms within the community
allogenic
environmental change governed by physical rather than biological processes.
3 elements of topography
altitude, slope, aspect
oxisols
old, highly developed, infertile rainforest soils
As soils develop
they get deeper, clay content increases, OM increases, CEC increases, horizons develop, pH decreases, water holding capacity increases
Oi
intact organic matter (ferric layer)
Oe
Partially fragmented organic matter (hemic layer)
Oa
highly decomposed organic matter (sapric layer)
Intermediate disturbance hypothesis
local species diversity is maximized when ecological disturbance is neither too rare nor too frequent
if disturbance is rare...
only the best competitors remain
if disturbance is frequent
only disturbance-tolerant species remain
Gross Primary Production (GPP)
total amount of CO2 fixed by plants
Net Primary Production (NPP)
GPP-Respiration
Respiration
total amount of CO2 lost by respiration
Exploitation Efficiency
how much of the resource will be consumed
Assimilation Efficienty
how much of the resource consumed will support growth and metabolism
Production Efficiency
Energy associated with growth
Essential elements are most available at pH of
6.5-6.9
What are life tables used for?
to describe age specific mortality as related to population growth rate. Constructed by following a population of individuals until they're all dead. Age specific mortality (qx)=individuals that died at age x/individuals alive at time x
sources of genetic variation upon which natural selection may act (4)
mutation, non-random mating, genetic drift, migration
how would increased stochasticity of mortality rate, survival rate, and birthrate affect the population?
Lambda (population growth) would vary more, wouldn't stabilize
what conditions favor exponential growth
large amounts of resources, lack of predators, lack of competition for resources, small or new populations
carrying capacity
maximum population size that an environment can support over time. occurs when b=d (lambda=0)
k=population of year x/population of year (x-1)
Why should managers care about natural selection
identify limited resources, increase fitness, increase genetic variation
adaptation
any characteristic of an organism that has evolved via natural selection that increases or at least maintains fitness under particular environmental conditions
natural selection
differential success of individuals within a population that results from their interaction with the environment
fitness
relative contribution of offspring to future generations
oxygen absorption
small organisms can easily absorb oxygen. Organisms with more cells may require other shapes or increased complexity ex: folding, branching
How does sphere size relate to SA:Volume ratio and what effect does that have on heat transfer
as sphere radius increases, SA:VOlume decreases. The more spherical something is, the less heat is loses to convection
optimal foraging theory
animals will tend t optimize calorie payoff for calorie ouput
marginal value theorem
organisms will hang out in a patch until there is a bigger energy benefit to go somewhere else
3 methods of predation
ambush, stalking, pursuit
Red Queen Hypothesis
There is an advantage for predators that are efficient at consuming prey.
There is an advantage for prey to adapt to evade predators
Predator chemical defenses
taste bad, look gross, or spray chemicals
cryptic coloration
camouflage
object resemblance
mimic the shape of a natural object
mimicry
one organism looks like another organism
aposematism
having intimidating colors which warn predators that they're dangerous
flashing coloration
distraction technique
armor
armor.
behavioral predator defense
ex: quails sit with their asses together in the nest to keep an eye out for predators
predator satiation
long life cycles with mass birth events
herbivore-biomass interaction
herbivores remove 6-10% of biomass annually from forests and 30-50% in grasslands
Impacts of herbivores on individual plants
loss of individual biomass, reduced growth rate, loss of nutrient reserves through selective tissue removal (fruit), loss of reproductive effort (buds/flowers/seeds)
3 herbivore defense strategies
structural, chemical, satiation
plant strucural defense
armor, unfavorable texture
plant chemical defenses
secondary compounds- nitrogen based compounds, terpenoids, and phenolics
nitrogen based compounds
deadly if consumed ex: morphine, caffeine, nicotine, cyanide
terpenoids
inhibit insects from reaching maturity/going through metamorphosis ex: oils, saps, latex
phenolics
smelly and indigestible ex: tannins, lignins
herbivore satiation
ex: "mast years" of the common beech tree. produce a huge amount of fruit. 3.1% of fruit is consumed during mast years while 38% of fruit is consumed most other years
intraspecific competition
competition between individuals of the same species
interspecific competition
competition between individuals of different species
6 types of interspecific competition
consumption, preemption, overgrowth, chemical reactions, encounter, territorial
consumption
individuals inhibit each other by competing for the same resource
preemption
beating other organisms to the resource
overgrowth
smother other organisms by growing on top of them or taller than them
chemical reactions
chemicals discourage other organisms from growing
territoriality
organisms defend their territories from other individuals
encounter
"flash territoriality" over a resource. One organism inhibits sharing of their resource.
competitive exculsion
lack of coexistence between competitors who occupy the same space and have the same needs
fundamental niche
all the environmental conditions under which a species could survive
realized niche
where a species is observed
parasitism
a symbiosis in which one organism (parasite) benefits from another (host), typically in a prolonged proximal and harmful interaction
intermediate host
parasite uses ahost for one or more developmental stages
definitive host
host in which a host reaches sexual maturity
parasitoid
a parasite that typically relies on an individual host and kills it
how can parasites impact hosts
increase host appetite, some parasites mimic growth hormones, impacts survival of hosts, can cause disease, can modify behavior, can make males look ugly so females don't mate with them, can cause organisms to lay more eggs so they have more hosts
2 host defenses against parasites
behavioral (remove ectoparasites manually),
inflammatory/immune responses (plants trap parasite eggs in shoots
mutualism
a kind of symbiosis in which both species benefit from their interactions
commensalism
one species benefits and the other isn't harmed
benefits of mutualism
transfer of nutrients, pollination, defense
mutualism & nutrient transfer
mycorrhizal bacteria fix Nitrogen
lichen=bacteria and fungus working together to get otherwise unreachable nutrients from rocks
Mutualism & defense
fish eat ectoparasites off sea turtles
ants live inside acacia tree stems. acacia fills stems with nectar. ants defend tree from predators.
mutualism & pollination
hummingbirds pollinate flowers as they consume nectar from flowers