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Phonemes
Unit of sound that forms the basis of speech and distinguishes one word from another in particular language
What is the IPA used for?
Differentiating sound and written word
Respiration
When air is pushed out of the lungs through trachea and up to larynx
Phonation
Vocal chords vibrate in response to air being pushed through them
More tension in vocal chords causes
Higher pitched voices, as do smaller vocal folds
Articulation
Process of producing speech via vocal tract and changes based on changes to jaw, lips, tongue
Formats
Resonant frequencies, peaks in speech spectrum
Three dimensions of spectrograms
Time, frequency, energy
Coarticulation
The overlap of articulatory or speech patterns, comes from experienced speakers positioning tongue before next vowels/constant
McGurk Effect
Resolves problem of coarticulation through vision, shows that we are affected by what we see
Motor theory of speech perception
Use reverse process of speech to understand what people are saying to us. We know how to speak and thus can infer what someone else is saying
Problem that arises from McGurk effect
speech production and vision perception is complex, so its limited
Solution to McGurk effect problem
Reduce the number of phonetic categories
Learning to Listen
Babies learn to distinguish particular acoustics early on before they start producing their own speech
What is a possible reason for the reduction in audible phonemes?
Forced categorization that may help us differentiate two sounds as different
Wernickes area
Responsible for associating sound patterns with meaning
Pathway for wenickes area
Primary auditoru cortex, Wernick, Broca, primary motor cortex
Brocas aphasia
Leads to difficulty in finding right articulation of words, able to understand what is being said to them
Wernickes aphasia
Production of coherent speech, inability to understand or assign meaning to sounds
Tone height and chroma
Height refers to level of pitch (related to frequency), chroma refers to shared qualities by tones that share an octave
Octave
Ration of 2:1, interval between 2 sound frequencies
What are semitones
12 equally spaced pitch intervals
Consonance
Pleasing combos of 2+ notes that arise from simple rations of fundamental frequencies
Dissonance
Unpleasant combos of 2+ notes that arise from complex rations of F.F, few coinciding harmonies
Scale
Particular subset of notes in a given octave
Key
Scale that functions as basis of musical composition
Tonic
Root notes of a key which acts as the gravity point of that key
What makes music pleasing?
Moving to and from root key
Where is music mostly processed in the brain?
Right auditory cortex and parabelt region
Melodic contour?
patterns tha rises and drops in pitch, defines melody regardless of the notes
Congenital amusia
Umbrella term for music deficiencies that arise not as a lack of intellectual ability, brain damage after brith or lack of exposure
Symptoms of amusia
Inability to distinguish small differences in pitch, tones that are out of key
ERAN and P600
Negative event related potentials occurring 200ms after melodic tonal violation. vs positive ERP that occur after 600ms
AMusics dont have P600 which suggests?
Lack conscious access to processed pitch deviances
Absolute Pitch
Rare ability where some people can very accurately name or produce notes without comparison to other notes
Proprioception and case study
Perception of body in space, based on ability to perceive the position of joints, includes verstibular sensations
Kinesthesia
Perception of movement w the exclusion of balance, uses the same receptors as proprioception
Muscle spindle
Kinesthetic nerve fibers attached to muscular fibers, perceives the stretching of muscles from within
Golgi tendon organ
in tendons, part of the muscle attached to articulation, able to perceive amount of pull in tendon
Tactile perception
Perception of objects that come into contact with the skin
Exteroceptive sense
perception of external world, associated with peripheral nerve fibers
Thermalgesia
Temperature and pain, uses interoceptive sense which is the perception of the internal state of the body
Somesthetic nerve fibers
difference in nerve fiber myelination between touch and proprioception versus pain fibers
Tactile perception
4 types, classified by receptive fields and rate of adaptation,
Slow adaptation rates will..
start firing when contacted with stimulus, and continue to fire as long as contact is maintained
Fast adaption will…
start firing as with contact to stimulus, stop firing if contact is maintained
SA1 Cells —> Merkel
Small receptive field with slow adaptation, good for fine grain patterns of tactile stimulation
SA2 cells —> Ruffini
Large receptive fields, slow adaptation, good for position of hands via skin
FA1 Fibers: Meissner
Large receptive fields, fast adaptation, good for maintain grip on objects
FA2 Fibers: Pacini
Fast adaptation rate, small receptive fields, good for interaction with tools
Example pathway of a key
Sa1 to feel the key → Sa2 to grasp key → Fa1 to maintain grip on key while unlocking door → Fa2 to feel when key has hit the end of the lock chamber
End sequence of touch fibers
converge to form nerve that enters the spinal cord between each spinal disc, patch of skin innervated by specific nerve called dermatome
Dosal column pathway
backside of spine, remains on the same side then crosses over onto 2 different nuclei, then to thalamus
types of nuclei
gracile and cuneate nucleus
Two point discrimination threshold
Perception of distance between two pins, higher number means larger distance between two pins to perceive pints as two distinct objects
Pain
an unpleasant sensory and emotional experience associated with or resembling association with an actual or potential tissue damage
Nociception
neural process of encoding nociceptive stimuli
Nociceptive stimulus
an actual or potentially tissue damaging event transduced and encoded by nociceptors
Two groups of peripheral nociceptors
A delta fibers and C fibers
A delta fibers
intermediate sized myelinated sensory nerve fibers that transmit pain and temperature signals
C fibers
small diameter non myelinated sensory nerve fibers that transmit pain and temperature signals
two stages that pain occurs in
sharp rapid pain followed by throbbing sensation
Throbbing sensation is mediate by…
c fibers
sharp rapid pain mediate by…
a delta fibers
how does local anesthetics work?
prevents sodium from entering sodium channel, this is how action potential propagates so no AP
Saltatory conduction
leads to jumping of AP from nodes of ranvier, thus 3 NoR rule to block AP in myelinated axons
Congenital insensitivity to pain
Prone to injury due to lack of nociceptive receptors
Limbic touch
Unmyelinated tactile afferents signal tough and project to insular cortex
For limbic touch, define both receptor types
Tactile receptors for sensation itself and receptors that encode pleasantness
Taste
perception of taste stimuli via interaction between soluble substances with gustatory receptors located in taste buds of the tongue
Smell
perception of smell using receptors located in olfactory mucosa in upper portion of nasal cavity
Trigeminal system
Chemesthesis is third chemosensory system next to smell and taste, receptors are located on fibers of the trigeminal nerve of the nasal and oral mucosa
What do trigeminal stimuli envoke?
irritation, burning or freshness
Taste buds located on
mucosa and papillae
Filiform papillae
somatosensory perception of the tongue, nothing to do with taste, texture of food
central pathway
tongue via cranial nerves to brainstem, to thalamus, insular and parietal cortex
2 anterior thirds are connect to brain via..
cranial nerve 7
posterior one third of tongue connected to brain via
cranial nerve 9
other taste buds found on epiglottis connected to brain via
cranial nerve 10
where do all cranial nerves connect to?
solitary nucleus of brainstem and then to the thalamus
Upper portion of nasal cavity contains
olfactory mucosa, sensory cells that are exposed to outside
Olfactory receptor neurons
Situated in roof portion of the nasal cavity, surrounded by supporting cells that regenerate stem cells
Olfactory code
humans have about 400 different types of olfactory receptors
specifications of olfactory receptor types
each ORN carry 1 receptor type, can be activated by different substances
Variation in human population
infinte types of combos for receptor types in each person, we all experience different smells
Where do all axons of the ORN pass through?
Cribiform plate of the ethmoid bone to reach the olfactory bulb
Olfactory bulb
where first synapses take place
Axons of olfactory neurons converge on one ….
glomerulus and do not cross over (occurs epsilaterally to brain)
Parts of olfactory processing areas that are part of the limbic system
amygdala, entorhinal, orbitofrontal cortex
Implications of direction to brain
memory and scene are intertwined, do not go through thalamus, create more vibrant memories triggered by scent
Orthonasal olfaction
volatile substances are released from the odor source and reach olfactory cleft within the respiration air stream
Retronasal smelling
substances reach odor source during swallowing
Smell functions
warning, nutrition, social communication
Anosmia
complete loss of olfactory function
Hyposmia
parietal loss of olfactory function
Parosmia
odors are perceived differently than they are supposed to
Phantosmia
perception of smells in the absence of an odor
Parkinsons and alzheimers
olfactory dysfunction predate diagnosis by 10-15 years
Factors that influence olfaction
age, sex, education level, bmi and blood preassure