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Homologous Structures
Structures derived from a common ancestor, may be modified for different functions
Examples of Homologous Structures
Human, cat, whale bat arms
Vestigal Structures
Remnants of ancestral (homologous) structures with no present adaptive function
Convergence Structures
Unrelated species have similar adaptions (analogous structures) under similar environmental conditions
Convergent Evolution
Natural selection acted in a way under some conditions (torpedo shape for swimming)
Allele
One of a number of alternate forms of a DNA sequence at a particular genetic locus
What has low mutation rates?
Plants and animals
Where do mutations happen faster and why?
In prokaryotes because they have short generation times
Crossovers, independent assortment, random fertilization
Sexual reproduction can result in genetic variation by recombining alleles
Microevolution
Change in allele frequencies in a population over generations
Popultion genetics
The study of what changes the allele frequencies in populations
Three mechanisms that cause allele frequency change
1.) Natural Selection
2.) Genetic Drift
3.) Gene Flow
Morphological Species Concept Definition
Organisms that have significant morphological & anatomical differences are different species
Example of Morphological Species Concept
Sorting birds into species based on their wingspan & beak size
Biological Species Concept Definition
Organisms that are reproductively isolated from each other are different species
Members of the same biological species
-Share the same gene pool
(there gene flow between species)
-Reproductively isolated from other species (by natural biological barriers)
Prezygotic Barriers
Act before the zygote is formed
Temperal Isolation
Species have different breeding schedules
Habitat Isolation
Members of a species move or are seperated
Behavioural Isolation
Actions or behaviours impacts reproduction
Gametic Isolation
Differences in gamete cells prevent fertilization
Postzygotic Barriers
Act after hybrid zygote is formed
Hybrid Inviability
An embryo is produced, but cannot survive development
Hybrid Sterility
Different species produce a viable offspring, but they can't reproduce
Hybrid
2nd generation hybrids are feeble or sterile
1.) Directional Selection
Selects phenotypes at one end of the spectrum of existing variation Shifts the populations genetic variance toward the new, fit phenotype
2.) Diversifying Selection
-Intermediates are less fit than extremes
-Maintains diversity
-Increases genetic variance
3.) Stabilizing Variation
Intermediate types more fit than extremes, Decreases genetic variation
4.) Frequency Variation
-The fitness of a phenotype depends on how common it is in the population
5.) Sexual Selection
Success based on traits related to obtaining mates (not directly related to the environment) Leads to sexual dimorphism
Intrasexual Selection
Individuals of one sex compete directly for mates of the opposite sex
Intersexual Selection
Individuals of one sex are choosy in selecting their mates from the other sex Typically female
Genetic Variation Definition
New genes and alleles can arise by mutation or gene duplication
Genetic Variation Facts
-Mutation rates low in animals and plants relatively
-Mutations accumulate quickly in prokaryotes & viruses because they have short generation times
-Only mutations in cells that produce gametes can be passed onto offspring
Who does natural selection act on?
Natural selection acts on individuals but a population evolves
What makes evolution possible?
Genetic variation in populations make evolution possible
Phenotypic Variation
-Phenotypic variation is mostly genetic
-The environment can influence expression, creating non-heritable variation
Mechanisms of Microevolution Definition
A change in allele frequencies in a population over generation
Three Mechanisms of Microevolution
1.) Natural Selection
2.) Genetic Drift (Founder effect & bottleneck effect)
3.) Gene Flow
Natural Selection- Mechanisms of Microevolution
-Acts non-randomly on the phenotypes of individuals
-Changes allelic & genotypic frequencies of populations non-randomly
-Always leads to adaption of population to current environment
Genetic Drift
-Genetic frequency changes due to randomness
-Often occurs in small populations (like sampling errors in statistics)
Outcomes of Genetic Drift
-Random changes in allele frequency in either direction
-Often reduces genetic diversity
-One allele may become "fixed" (all other alleles lost)
The Founder Effect & Genetic Drift Definition
-A few founders start a new isolated population
The Founder Effect & Genetic Drift Facts
Founder gene pool differs from isolated source, Small population size leads to more drift, Better alleles may be lost
The Bottleneck Effect & Genetic Drift
An event drastically cuts population size
Gene Flow- Mechanisms of Microevolution
Alleles move in/out of a population
Gene flow Includes
-Migration of adults
-Dispersal of gametes, seeds & larvae
Relative Fitness Definition
Fitness is relative to other individuals in the population
Fitness Definition
Fittest= best reproductive success
Hardy Weinburg Equilibrium Definition
If a large population reproduces sexually at random, then the genetic frequencies should not change in next generation (remains in equilibrium)
Hardy Weinburg Conditions
1.) No mutations
2.) Mating is random
3.) No selection (equal survival)
4.) Very large population size
5.) No gene flow in or out
What does Hardy Weinburg let us detect?
-Microevolution
-If actual ratios ≠ expected, population is evolving
-Microevolution: an evolving population is one that is showing genetic change over generations
Phylogeny Definition
-The evolutionary history of a species & its relationship to other species
-Shown as a "tree"
Types of Phylogentic Trees
rooted, unrooted
Rooted Phylogenetic Trees
Single lineage (at base) represents common ancestor
Unrooted Phylogenetic Trees
Shows relationships but not a common ancestor
Cladistic Analysis Definition
looking at the clades and seeing similar traits
Monophyletic Group (Clade)
Consists of an ancestral species and all of its descendants
Paraphyletic Group
Consists of ancestral species and some, but not all, of its descendants
Polyphyletic Group
Includes distantly related species but does not include their most recent common ancestor
Endosymbiont Definition
A cell which lives inside another cell with mutual benefit
How Did eukaryotic Cells Evolve
-Eukaryotic cells are believed to have evolved from early prokaryotes that were engulfed by phagocytosis
-The engulfed prokaryotic cell remain undigested as it contributed new functionality to the engulfing cell (e.g photosynthesis)
-Over generations, the engulfed cell lost some of its independent utility and became a supplemental organelle
What Organelles Came Via Endosymbiosis
Mitochondria & chloroplasts are both organelles suggested to have arisen by endosymbiosis
Genome Fusion
-The eukaryotic nucleus resulted from fusing archaeal & bacterial genomes
-Gram negative bacteria, which have two membranes, resulted from fusing Archae & Gram-positive bacteria, each of which has a singular membrane
What are the two types of bacteriophage life cycles?
Lytic and lysogenic
What happens in the lytic phage life cycle?
The phage lyses the host cell after replication of their virion, releasing phage progeny to find new hosts
What happens in the lysogenic phage life cycle?
The phage genome integrates with the host genome, replicating without destroying the cell; the phages are known as temperate phages
What is a prophage?
When lysogenic phage DNA is incorporated into the host cell genome
When do lysogenic phages initiate the lytic cycle?
When conditions deteriorate for the host cell, such as a lack of nutrients
What is the first step of the lytic cycle?
Phage infects cell
What happens to the phage DNA in the lytic cycle?
Phage DNA circularizes, remains separate from host DNA
What occurs during the third step of the lytic cycle?
Phage DNA replicates & phage proteins are made, new phage particles are assembled
What is the final step of the lytic cycle?
Cell lyses, releasing phage
What is the first step of the lysogenic cycle?
Phage infects cell
What happens after phage DNA becomes incorporated into the host genome in the lysogenic cycle?
The cell divides and prophage DNA is passed to daughter cells
What triggers the excision of phage DNA from the bacterial chromosome in the lysogenic cycle?
Under stressful conditions
What occurs after the phage DNA is excised from the bacterial chromosome in the lysogenic cycle?
Phage DNA replicates & phage proteins are made, new phage particles are assembled
What is the final step of the lysogenic cycle?
The cell lyses, releases phages
When did the first organisms on Earth appear?
3.5-3.8 billion years ago
How many species of prokaryotes exist?
Millions
Where can prokaryotes be found?
Everywhere, on & in every other living organism
What role do prokaryotes play in the ecosystem?
They are essential to all life, often benign
The Ancient Atmosphere & Prokaryotes
Anoxic Definition
No molecular oxygen
Phototrophs
Autotrophic organisms that convert solar energy into chemical energy
When Did Phototrophs First Appear
Within one billion years of the formation of the earth
Cyanobacteria (Blue-Green Algae)
Evolved from simple phototrophs at least one billions years
Ancestral Cyanobacteria
-Began "oxygenation" of the atmosphere
-Increases in O2 concentrations allowed the evolution of other life forms
Prokaryotic Cell Size & Structures
-Prokaryotic cells are significantly smaller than eukaryotic cells
-Predominantly single celled organisms of the domains bacteria & archaea are prokaryotes
All Cells Have Four Common Structures
1.) The plasma membrane
2.) The cytoplasm
3.) Double stranded DNA genomes
4.) Ribosomes
The Plasma Membrane
Functions as a barrier for the cell & separates from its environment
The Cytoplasm
Complex solution of organic molecules that salts the inside of the cell
Double Stranded DNA Genome
Informational archive of the cell
Ribosomes
Sites of protein synthesis
Gram Positive Bacteria Definition
-Includes many pathogens
-Many decomposers in soil
(Ex: Bacillus anthracis (anthrax) & Clastridium tetani (tetanus))
Gram Stain Reflects Cell Wall Type (+)
-Gram + bacteria with thick PG layer
-Stains purple in gram stain
Gram Negative Bacteria
-Bacteria with thinner PG layer plus outer lipid bilayer membrane
Gram Stain Reflects Cell Wall Type (-)
-Pink in gram stain; outer lipopolysaccharide layer often toxic, resists drugs & immune system
Endotoxins
Toxic outer membranes of some gram negative bacteria,Generalized toxic effect (Salmonella; food poisoning)
What are endospores?
Endospores are dormant, non-reproductive structures produced by some bacteria under stress.