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Topics: 29 - Fungi 33 - Viruses 34 - Plant From and Function 35 - Water and Sugar Transport in plants
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Organelles
originated from more primitive cells that become trapped in eukaryotic cells
Fungi
eukaryotes
single celled or multicellular filaments
most important decomposers
are mutualists
positive impacts of fungi
course of antibiotics
food
yeast for bread, cheese, soy sauce, beer, wine
improve characteristics of foods such as fruit juice, candy, and meat
mycorrhizal fungi
live close to plant roots
plants grow better in their presence
name for mycorrhizal fungi
mycorrhizae
saprophytes
fungi that digest dead plant material
cycle carbons through terrestrial systems
what do fungi do for the carbon cycle?
speed it up
cell wall
rigid
structural support and shape
made of chitin
cytoplasmic membrane
bilayer of phospholipids and protein molecules
contain sterols
reduces membrane fluidity and permeability
selective permeable barriers
controls what comes in and out of cell
two growth forms of fungi
single cell form (yeast)
multicellular, filamentous forms (mycelia or mycelium)
nature of mycelium
dynamic
grow in direction of food sources and die back in areas where food is running out
body shape can continuously change
Hyphae
long, narrow filaments of mycelium
septa
cross walls in hyphae
pores allows materials to flow between compartments
coenocytic
lacking septa (cross walls)
many nuclei are scattered throughout the mycelium
mycelia supports what type of digestion
external
swimming gametes and spores
sexually produced gametes and asexually produced spores (zoospores) or chytrids have flagella
zygosporangia
spore producing structures of zygomycetes
form from fusion of cells from joined together haploid hyphae from two individuals
basidia
basidiomycete (club fungi) form basidia
ends of hyphae
produces 4 spores
Asci
ascomycetes (sac fungi) form asci, reproductive sac like cells at ends of hyphae
produce 8 spores
conidia
asexual spores
dispersed by water or wind currents and grow into new hyphae
why are fungal infections in humans more difficult to treat?
similar cellular and molecular structures
drugs that disrupt fungal physiology are likely to damage humans as well
symbiotic relationship
living together (between plant and fungi)
mutualic relationship
benefit both species
parasitic relationship
one benefits other is harmed
commensal relationships
benefit one another is unaffected
types of plant mycorrhizal interactions
ectomycorrhizal fungi (EMF)
arbuscular mycorrhizal fungi (AMF)
Ectomycorrhizal Fungi (EMF)
found in tree species in temperate regions
form a dense network of hyphae that cover plant roots
enter soil but not root cells
penetrate decaying material
release peptidases
provide phosphate ions to host plant
peptidase
cleave proteins to release amino acids that hyphae transport
Arbuscular Mycorrhizal Fungi (AMF)
hyphae grows into the cells of root tissue
also called endomycorrhizal fungi
endophytes (inside plant)
organisms that live between and within plant cells
live in close association with roots or aboveground tissues of land plants
Lichens
mutualistic partnership usually between a specie of ascomycete and either a cyanobacterium or an alga
extracellular digestion
digestion that takes place outside the organism
simple compounds resulting from enzymatic action are absorbed by hyphae
what aids in lignin degradation?
enzyme called lignin peroxidase
what aids in cellulose digestion?
cellulases
what are the most fundamental reproductive cell in fungi?
spores
what do mating types also function as?
sexes
how does fertilization occur in fungi?
fusion or hyphae or fusion of nuclei from the fused hyphae
plasmogamy
occurs when the cytoplasm fuse
heterokaryotic
when two or more genetically distinct nuclei exist within a single mycelium
what are most heterokaryotic hyphae?
dikaryotic
karyogamy
occurs when the nuclei fuse to form a diploid zygote
cyrtid life cycle
alternation of generations
Key stages of the chytrid life cycle:
1. Haploid adults form gametangia: ▪ Mitosis produces male and female swimming gametes
2. Gametes fuse to form a diploid zygote
3. The zygote grows into a diploid sporophyte
4. Haploid spores, which disperse by swimming, are produced by meiosis inside the sporophyte’s sporangium
Zygomycete Life Cycle
hyphae from different mating types fuse
can also reproduce asexually by making sporangia, which produce haploid spores by mitosis
Basidiomycete Life Cycle
Mushrooms are sexual reproductive structures produced by basidiomycetes
originate from the dikaryotic hyphae of mated individuals:
Ascomycete Life Cycle
Hyphae or specialized structures from different mating types fuse:
Forms a heterokaryotic
Short dikaryotic hypha with cells containing two nuclei emerges:
asci at tips
production of eight haploid spores • When the ascus matures, the spores inside are forcibly ejected
Microsporidia
Single-celled and parasitic
polar tube
Chytrids
Largely aquatic and common in freshwater
digesting cellulose:
important decomposers of plant material in wet soils, ponds, and lakes
Many freshwater species are parasitic: – Likely culprit behind the decline of amphibian populations worldwide
Zygomycetes
Primarily soil-dwellers
responsible for rotting fruits and vegetables
Glomeromycota
All members (except for a single species) comprise the arbuscular mycorrhizal fungi (AMF)
Absorb phosphorus, nitrogen, other nutrients, and water into roots of most of plants living in grasslands and tropical forests
Host plant provides symbiotic fungi with sugars and other organic compounds
Basidiomycota
Form mycelia and produce basidia where meiosis and spore formation occur
Mushrooms
wood decomposition Some are ectomycorrhizal fungi (EMF) that improve the health of forests Some can also be parasitic
Ascomycota
Most form mycelia and produce spores in asci: – Mycelia also commonly reproduce asexually • yeasts
mutualistic association with cyanobacteria and/or single-celled green algae, forming structures called lichens
mutualistic EMF associations with tree roots
properties of viruses
not cells
active only inside host cell
multiply by taking control of host
lack enzymes for most metabolic processes
lack machinery for synthesizing proteins
contains a nucleic acid core
capsids
shell surrounds the nucleic acid
Nucleocapsid
capsid and nucleic acid together
Envelope
• Not found in all viruses
• Usually a modified piece of the host cell membrane
Spikes
• Found on both naked and enveloped viruses. Project from either the nucleocapsid or envelope
• Allow viruses to dock with their host cells
Virion
Fully formed virus able to establish infection in a host
Capsomeres
Identical protein subunits that spontaneously self assemble to form the capsid
Helical capsid
Rod-shaped capsomeres that form a continuous helix around the nucleic acid
Icosahedral capsid
Three-dimensional, 20-sided figure with 12 evenly spaced corners
Complex capsids
Found in bacteriophage, the viruses that infect bacteria
Have multiple types of proteins
Take shapes that are not symmetrical
Viral Envelope
• Composed of the membrane system of the host • Cell membrane or nuclear membrane
• Regular membrane proteins are replaced with viral proteins
• Spikes: protruding glycoproteins essential for attachment to the host cell
what type of genetic material can a virus have?
DNA or RNA but not both
general phases in the life cycle of animal viruses:
Adsorption
• Penetration
• Uncoating
• Synthesis
• Assembly
• Release from the host cell
Adsorption
Invasion begins when the virus encounters a susceptible host and adsorbs specifically to receptor sites on the cell membrane
•Endocytosis
entire virus is engulfed by the cell and enclosed in a vacuole or vesicle
Uncoating
enzymes in the vacuole dissolve the envelope and capsid, releasing the virus into the cytoplasm
where do RNA viruses replicate?
cytoplasm
where do DNA viruses replicate?
nucleus
exocytosis
Enveloped viruses are liberated by budding
Nucleocapsid binds to the membrane
• A small pouch is formed
• Pinching off of the pouch releases the virus with its envelope
• Viruses are shed gradually without destruction of the cell
Inclusion bodies
compacted masses of viruses or damaged cell organelles in the nucleus or cytoplasm
Syncytia
fusion of multiple host cells into single large cells containing multiple nuclei
Oncogenic viruses
Experts estimate that up to 13% of human cancers are caused by viruses
Transformation
Virus carries genes that directly cause cancer
• Virus produces proteins that induce a loss of growth regulation in the cell
Viruses That Infect Bacteria
bacteriophage
Lytic phase or lytic cycle
life cycle of bacteriophage that ends in destruction of the bacterial cell
Lysogenic cycle
bacteriophage becomes incorporated into the host cell DNA
Temperate phages
Have the ability to undergo adsorption and penetration but do not immediately undergo replication or release
Prophage
an inactive state in which phage DNA is inserted into the host chromosome
Induction
activation of a prophage in a lysogenic cell to progress directly into viral replication and the lytic cycle
lysogenic conversion
The acquisition of a new trait from a temperate phage
Responsible for the diphtheria toxin, cholera toxin, and botulism toxin
•Interferon (IFN)
–Naturally occurring human cell product
–Used with some success in preventing and treating viral infections
Viroid
Virus like agent that parasitizes plants
Composed only of naked strands of RNA—lack a capsid or other type of coating
what type of growth do most plants undergo?
indeterminate growth
Aboveground: – Shoot system
harvests light and carbon dioxide from atmosphere to produce sugars
Belowground: – Root system
anchors plant and takes in water and nutrients from soil
Root and shoot systems both function in what?
absorption
key functions of the root system:
– Roots anchor plant to the soil
– Roots absorb ions and water from soil
– Roots conduct water and ions to the shoot system
– Roots obtain energy from the sugar in the shoot system
– Roots store material produced in the shoot system for later use
Morphological Diversity in Root Systems
herbaceous
perennial
herbaceous
Seed plants that lack woody tissue
perennial
They live for many years
Phenotypic Plasticity in Root Systems
– Their form is changeable depending on environment
Modified Roots
formed by various types of plants
Adventitious roots
grow from shoot system: – In ivy, adventitious roots act to anchor the plant
Prop roots
brace the plant (corn)
tap root
originates from radicle
Pneumatophores
specialized lateral roots in genus Avicennia
Function in gas exchange
Oxygen in atmosphere can diffuse into root system through pneumatophores
storage roots
Thick taproot that stores carbohydrates during plant’s first two growing seasons
• Biennial plants, such as carrots and beets: