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colonial/modular organisms
individuals are genetically identical
if colonial organisms are connected, they compose a module
Modular organisms are brain coral, soft coral, byrozoan
gonochoristic
have separate sexes
different than sexually dimorphic
sequential hermaphrodites
an organism is one sex for a time and then another sex
protandrous - male then female
protogynous - female then male
simultaneous hermaphrodites
organs to produce male and female gametes present simultaneously (not necessarily active simultaneously)
size advantage model
for protandry
eggs are costly, so more offspring produced when individual functions as female when large
male reproductive function does increase with size, therefore there is a threshold size when female improves
smaller individuals do better as males
male polymorphism
males may occur as aggressive fighting morph or less aggressive morphs
morph determination can be environmental or genetic
less aggressive morphs can obtain mates by “sneaky” tactics, which are often successful
diadromous
move between estuaries and the open sea for migratoin
andadromous
fish live as adults in salt water, spawn in freshwater (shad, striped bass), more common in higher latitudes
catadromous
fish live as adults in freshwater, spawn in salt water (eel), more common in lower latitudes
oceanodromic
fully oceanic (herring, green turtle, humpback whale)
planktotrophic dispersal
female produces many (10³-10^6) small eggs
larvae feed on plankton
long dispersal time (weeks)
some are very long distance (teleplanic) larvae that cross oceans
lecithotrophic dispersal
female produces fewer eggs (10²-10³)
larger, larvae live on yolk
short dispersal time (hours-days usually)
what is the benefit of sexual reproduction?
crossing over and mixing of genes produces more genetic diversity and increases survival of individuals
what is a parasitic male and where would you find one? what is the benefit of this?
parasitic male latches onto female and atrophies until he dies and sperm would be available for female
you find these in the deep sea
the benefit is that you don’t have to search for a mate in the deep sea where organisms are few and far between
how do dispersal and migration differ?
dispersal is undirected
migration is directed, movement between specific sites, why specific areas? why the route and length?
compare and contrast planktotrophic and lecitotrophic dispersal mechanisms?
planktotrophic has long dispersal time and distance, larvae feed on plankton
lecithotrophic larvae live on yolk and have a short dispersal time
what can larvae control when it comes to dispersal?
larvae can control time of larval life (competency, overall development), timing of upward and downward movement, final micro-movements to “good” adult habitat
why might an organism want/need to disperse?
avoidance of crowding, local extinction, hedging bets (spread over habitats)
Compare and contrast protandrous and protogynous hermaphrodites. Give examples. Why might an organism adapt this reproductive strategy?
protandrous are male then female (clownfish) - large females can make a lot of eggs, small males compete less
protogynous are female then male (grouper) - large males can defend territory and make a lot of eggs
pick an organism that migrates and be able to describe why it migrates, where it migrates, and what benefits migration provides
sockeye salmon migrate from the ocean to specific rivers and pools where they were born to spawn, this benefit allows them to return to safe breeding locations year after year and be sure that other salmon are in the area
plankton
organisms living in the water column, too small to be able to swim counter to typical ocean currents
holoplankton
spends entire life as plankton
meroplankton
part of life spent as plankton
cyanobacteria
blue green algae
most abundant photosynthetic organism
picoplankton (1 micrometer)
nitrogen fixation
nematocysts
stinging or sticky grabbing cells - on tentacles
trait of Cnidaria
phytoplankton
diatoms, dinoflagellates, cyanobacteria, coccolithophores
photosynthetic, live in water column but too small to swim countercurrent to typical ocean currents
zooplankton
plankton that eat other animals - can’t swim against ocean currents
briefly describe diatom reproduction
they reproduce asexually by binary fission (also use sexual reproduction)
there is a small and large side of their frustule (silica skeleton) so they will split and each form a new diatom
get smaller and smaller until they are too small, then they have to sexually reproduce
briefly describe nitrogen fixation
cyanobacteria do this
converts gaseous nitrogen into NH4+
available for synthesis of amino acids and proteins
compare and contrast diatoms and dinoflagellates
compare - they both asexually and sexually reproduce, they have some sort of outer casing
contrast - diatoms occur singly or form chains, do not swim, usually radially symmetrical, reproduce very fase
dinoflagellates - have two flagellae and can swim, have several life history stages such as benthic cysts, heterotrophic, abundant in tropics, mid-latitudes in summer, can cause red tides
briefly explain how algal blooms occur and why do we care?
algal blooms occur when there is an influx of nutrients, mainly nitrogen and phosphorus, that increase the carrying capacity of algae which reproduce out of control
we care because it disrupts ecosystems, creates hypoxic zones, and creates harmful neurotoxins in the water
compare and contrast cnidarians and ctenophores
cnidarians are scyphozoans and siphonophores, they are mainly carnivores, and use nematocysts to hunt, siphonophores can be colonial
ctenophores have combs which refract and flash light (Newton rings), micro carnivores with two long sticky tentacles and feed on smaller zooplankton, planktonic eggs, invertebrate larvae, and bioluminesce with luciferin-luciferase system
briefly describe the importance of crustacean zooplankton
main food of many megafauna such as baleen whales
nekton
organisms that can swim against current
otolith
calcium carbonate ear stones that sense vibrations which travel to auditory brain region for processing
lateral line
excellent mechanoreception, line down body which can sense movement in water
pinniped
seals, sea lions, walruses
mustelidae
sea otters
sirenian
sea cows, dugongs
Describe the functional-morphology plane for locomotion in fishes (distinguish between acceleration, cruising, and maneuvering specialists
acceleration specialists have minimum resistance (barracuda)
maneuvering specialists have paired fins around center of mass for steering and stabilization (buttterfly fish)
cruising specialists minimize drag and have a high aspect ratio tail with stiff streamlined body (tuna)
what is the difference in swim bladder rete mirable lengths between shallow and deep sea fish?
deep water fish have longer rete mirable than shallow water fish because they can increase the maximum air concentrations in their swim bladder to cope with high pressure in deep water
what are the three different feeding mechanisms fish use?
ram feeding (overtake prey by ramming water through mouth (filter feeding))
suction feeding (create suction by rapid expansion of buccal cavity)
manipulation (biting, scraping, grasping, usually with use of teeth)
why do fish need to hear?
to avoid predators, communicate, and water is effective at transmitting sound
what is the biggest difference between Odontoceti and Mysteceti?
Odontoceti are toothed whales
mysticeti are baleen whales
describe some key features of seabirds
often colonial breeders, monogamous, courtship involves elaborate displays, crowded breeding sites with several species, protected from predators, feeding involves either diving or underwater swimming, long distance migration between nesting and feeding areas is common
what are the two problems with diving in marine mammals? how do marine mammals achieve such long dives (name at least 2-3 reasons why)
problems - must breathe at surface and have oxygen for long dives
they do this by increased volume of arteries and veins, increased blood cell concentration, decrease heart beat rate and O2 concentration, can restrict peripheral circulation and circulation to abdominal organs
describe some key features of the sea turtle life cycle
females return to same beach, eggs hatch a few weeks after being laid, hatchlings crawl toward shoreline using cues of light and magnetic field of the earth
predation on hatchlings is very high
adults use magnetic field of earth
sex determination is temperature dependent, females are born in colder temperatures and males are born in warmer temperatures
diurnal vertical migration
zooplankton rise to shallow water at night and sink to deeper water during the day
start to sink before dawn, rise before dusk
internal biological clock that is reinforced by day-night light changes
predation hypothesis
cause of vertical migration is based on visual prey detection
counterillumination
fish have ventral bioluminescent organs, camoflages them from below (deep sea organisms)
bioluminescence
either luciferin-luciferase or photoprotein-calcium system to avoid predation or help hunt
what are some of the defenses plankton utilize?
large elaborate spines, transparent bodies, toxic substances, bioluminescence
what are the benefits of schooling?
protect against predators (spherical aggregation, have trouble attacking individual fish, many eyes theory), reduction of drag (slipstreaming fish in front, fish in front exchange places periodically)
describe what happens during the diurnal vertical migration. describe one of the hypotheses used to describe the process
strong light hypothesis (zooplankton are adversely affected by UV radiation)
phytoplankton recovery hypothesis (zooplankton exploit phytoplankton but allow them to recover)
surface mixing hypothesis (surface waters bring more food when zooplankton return)
energy conservation hypothesis (lower metabolic cost in cooler water)
limiting nutrients
nitrogen, phosphorus, silicon, iron, trace elements
briefly describe the nitrogen cycle
gaseous N2 from atmosphere to nitrogen fixation (NH3) to organic/inorganic nitrogen then denitrification (back to N2) back to atmosphere
be able to describe what is happening in Figure 11.1
over the winter there are high phosphates and nitrates at the surface of the water and available sunlight increases
in the spring as sunlight availability increases there is a large spring diatom bloom with a zooplankton bloom lagging behind
as they consume all the nutrients there is a bust, and then as nutrients accumulate there is a second smaller diatom bloom
which nutrient is generally thought to be most limiting and what are some sources of it?
nitrogen is believed to be the main limiting element for phytoplankton growth - sources include nitrogen cycle and runoff from human development
what would happen if more of the limiting nutrient were added to the ecosystem? what type of ecological regulation would this be an example of?
the phytoplankton population would increase
bottom up regulation
biomass
mass of living material present at any time
g/unit area or volume
productivity (primary and secondary)
rate of production of living material per unit time per unit area/volume
primary is due to photosynthesis, secondary is due to consumers of primary producers
food chain
linear sequence showing which organisms consume which other organisms, making a series of trophic levels
food web
more complex diagram showing feeding relationships among organisms, not restricted to a linear hierarchy
gross primary productivity (GPP)
total carbon fixed during photosynthesis
Net Primary Productivity (NPP)
total carbon fixed during photosynthesis minus the part which is respired
Why is energy lost from one trophic level to the next (three reasons)
some material not eaten, not all food eaten is converted with 100% efficiency, metabolic costs are a loss
describe three reasons for the differences in geographic variation of productivity
continental shelf and open-ocean upwelling areas are most productive
convergences and fronts often are sites of rise of nutrient rich deep waters
central ocean, gyre centers are nutrient poor, low primary production