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complex systems
new genes evolve new functions
old genes take on new functions
genetic toolkit
different animal body plans are built with the same basic
homeotic transformations
one body part replace another
certain mutations most common in parts of the body that were either repeated
evo-devo
variations in structures depend in part on when and where certain genes are expressed
also called molecular paleontology
homeotic genes *
control foundation of tissues
act as signals that create a sort of instructional map for where structures should develop
homeotic genes cont.
affects phenotype and is which natural selection acts
hox genes *
homeotic genes in animals
position on chromosome correlates to the relative position on the animal body part that it regulates
deep homologies *
genes responsible for prominent and widely divergent traits come from a common ancestor
pax6 is an example (eye sensor)
photoreceptors
captures light
ancestors of opsins
receptor for the hormone melatonin
act as sensors grabbing molecules passing by and relating signal inside the cell
evolution is tinkering not engineering *
does not improve organisms to perfection
well enough traits will be selected
phenotypes are results of trade offs and physical constraints
trade-offs *
birds: trade off between total field and range of binocular vision
owl (high binocular low monocular)
costly
sexual reproduction is
benefits of sex *
speeds up pace of adaptation
cannot separate bad mutation from good in asexual reproduction
sexual combines genotypes in each generation into new combinations
Muller ratchet *
purges deleterious mutations - sexually produced offspring are genetically different from their parents
recombination creates two new variants, which reduces deleterious mutations
Red queen hypothesis *
allow rapid evolution response to parasites predators
Fisher-muller hypothesis
generates genetic variation that is favored by natural selection accelerates adaptive evolution
bring 2 beneficial mutations together
sexual dimorphism *
systematic difference in form between individuals of different sex in the same species
suggests sexual selection affects organisms differently
ie: big spider female, and small male
persuades members to mate
sexual selection *
traits that better help attract other members to reproduce
female parental investment *
increase relative fitness of young receiving
reduces reproductive potential of parent giving it
intersexual selection*
males showw off color, behaviors; females choose the most attractive
intrasexual selection*
male to male competition and female have no right to choose
female choice drives evolution and showy traits
traits become exaggerated
trade-offs
ie: sperm battles
sexual conflict
increases fitness of male and lowers reproductive success of females
species *
fundamental units of organism diversity
species diversity
half known species are insects
recognizing species
survival depends on our ability to recognize species
origin of species*
direct result of evolution
biological species concept
useful due to focusing on the process of how species form
hybrid problem still exist
allopatry speciation*
geographical barriers preventing from interbreeding
physical separated
parapatric speciation*
biological barriers
food preferences, spawn time etc
occurs in a hybrid zone
sympatric speciation*
species diverge while inhabiting the same physical space
resource competition and reproductive isolation
hybrid zone*
areas of contact between 2 diverging population where hybrids are common
disapproves when speciation is complete
reinforcement*
when NS acts to increase the amount of reproductive isolation between population
ring species*
population at both ends of a range grow more different from each other due to some factors (cannot interbreed due to genetic divergence and therefore leading to reproductive isolation)
phylogenetic species concept*
2 organisms with different evolution history are different species
cons: requires comprehensive analyses
pros: testable, objective and applicable
macroevolution
history of biodiversity
process of organization, adptation, and extinction
biodiversity
increases if rate of origination increase
increases if rate of extinction decrease
biogeography*
study of how continental drift affect organisms
vicariance*
physical barriers affecting organisms overtime
punctuated equilibrium*
evolution is not slow and steady
can be small bursts often associated speciation event
adaptive radiations*
dramatic bursts of evolution and lead to convergent evolution
endemic species - found only in a certain area due to the particular niche
three groups of fauna
cambrian - both high origination and extinction rates
paleozoic - lower origination and extinction rates
modern - lowest origination and extinction
the big five mass extinction events*
permian and triassic 252 mya (96 percent were gone)
could be caused by volcano eruption
most recent is 66 mya caused by asteroid
dinosaurs*
not extinct, but birds now
coevolution*
the reciprocal evolutionary change between ecologically intimate species
one species may cause selection to operate in new ways on the other
mutualistic interactions*
both species benefit from the interaction
must involve substantial genetic modifications in both partners
antagonistic coevolution*
evolutionary changes in each species decreases the fitness of the other species
cospeciation
speciation in one species leads to speciation in the other
caused by high benefits and long term mutualistic relationship
obligate mutualism
each partner can only survive and reproduce successfully in the presence of the other
often leads to cospeciation
cheaters
reap the benefits of cooperation without having to pay the associated costs
antagonistic coevolution*
predator prey and parasite host e
evolution arms race*
evolve at the same tiem to survive
mimicry*
one resemble another so predator is confused
aposematic
warning coloration
batesian*
share looks but no defensive attribution
mullerian*
share looks with defensive attribution
coevolution
___leads to dependence and can lead to extinction
cultural transmission*
when learning from others, information spreads through the population quickly
cultural evolution
when cultural transmission leads to changes in the frequency of traits