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How many Extant Fish Species
Approximately 35,800
Epipelagic Zone
Ocean zone from the surface to 200m.
Mesopelagic Zone
Ocean zone from 200m to 1,000m.
Bathypelagic Zone
Ocean zone from 1,000m to 4,000m.
Abyssopelagic Zone
Ocean zone from 4,000m to 6,000m.
Hadalpelagic Zone
Ocean zone below 6,000m.
What does hydrostatic pressure effect at deep depths
Protein formation.
Tri-methylamine N-oxide (TMAO)
Stabilises proteins in fish tissues, concentration increases with depth.
Paraphyletic
Common ancestor in group but not all descendents are in the group.
Agnatha
Jawless vertebrates.
Class Petromyzontida
Lampreys; most extinct, 48 extant spp.
Class Myxini
Hagfish; most extinct, 91 extant spp.
Anadromy (anadromous)
Adult life in sea, returning to freshwater to breed.
Diadromy (diadromous)
Migration between freshwater and seawater. The principal feeding and growing biome differs from the reproductive biome.
Catadromy (catadromous)
Fish that live in freshwater and return to the sea to breed.
Gnathostome
Jaws
Mandibular arch
Becomes biting jaws
Class Chondrichthyes
Cartilaginous fish, including sharks, skates, rays, and chimaeras.
Sub-class Elasmobranchii
Sharks, Skates, Rays (1,283* extant spp.)
Class Osteichthyes
Bony Fish (34,344* extant spp.)
Chondrichthyes Characteristics
Cartilaginous fish, Spiracle, No operculum opening, No swim bladder, Ampullae of Lorenzini, Spiral valve, Placoid scales (denticles), Internal fertilization.
Osteichthyes Characteristics
Bony Fish, No spiracle, Operculum covers gills, Swim bladder, No electro receptors, No spiral valve but pyloric caecae, Overlapping scales, Mostly broadcast spawners.
Lower teleosts Characteristics
Soft fin rays, 1 dorsal fin (maybe adipose), Pectoral fins low, Pectoral fin base horizontal, Pelvic fins abdominal, Pelvic fins with numerous rays, Scales usually cycloid, Swimbladder often physostomatous
Higher teleosts Characteristics
May also have spiny fin rays, 1 or more dorsal fins, Pectoral fins higher, Pectoral fin base angled towards vertical, Pelvic fins thoracic or jugular, Pelvics fins with reduced number of rays, Scales usually ctenoid, Swimbladder physoclistous
What percent of all fish are teleosts
96%
What are the “big seven” fish families by species count?
Cyprinidae, Gobiidae, Characidae, Cichlidae, Labridae, Loricariidae, Serranidae
What is the largest vertebrate order?
Perciformes
Which family is known for fused pelvic fins forming an adhesive disc?
Gobiidae
What is a “species flock”?
A group of closely related species derived from a common ancestor, often in isolated habitats like African Great Lakes.
what is the Characidae family
characins, piranha
What is the Cyprinidae family
minnows, carps - largest fish family
what is the Gobiddae family
gobies
what is the Cichildae family
cichlids
what is the Labridae family
Wrasses
what is the Loricariidae family
armoured catfish
what is the family Serranidae
groupers
What are clupieformes
herrings, sardines, shads, menhaden, anchovies - very important commercially
What are Gadiformes
cod, haddock, pollock, whiting - very important commercially
what is a physostome
a type of fish with a swim bladder connected to the gut, allowing for gulping air.
what is a physoclist
a type of fish with a closed swim bladder that is not connected to the gut, regulating buoyancy through gas exchange.
What is the Bohr effect
Describes a reduction in haemoglobin-oxygen affinity under lower pH
what is the Root effect
Describes a reduction in the oxygen carrying capacity of hemoglobin at low pH, specifically in some fish species.
what is the primary function of the caudal fin
provides thrust and propulsion for swimming.
what is the difference between median and paired fins
Median fins - Dorsal, aal, caudal
Paired fins - pectoral, pelvic
what tail is best for short bursts of speed and has high drag
low aspect ratio tails. e.g. rounded, truncate (sole, flatfish)
what type of tail is best sustained for swimming and has low drag
high aspect ration tails. e.g. forked lunate (tuna)
what is the aspect ration of a tail
Aspect Ratio = (Tail height) squared / Tail area
what is a myotome
a segment of axial muscle (contracts to generate flexion)
what is the function of red muscle
slow, continuous swimming (long durations)
what is the function of white muscle
Fast, powerful bursts (used briefly)
What is Body Caudal Fin Swimming (BCF)
swimming powered by side to side movements
What is Labriform swimming
Swimming using pectoral Fins (wrasses, parrot fish)
what’s the ideal body shape for continuous swimming
Fusiform (streamlined)
what do fillets in scombroids do
improve hydrodynamic efficiency
Which fin types help prevent pitch, roll and yaw
pitch - pectorals and pelvics
roll and yaw - dorsal and anal
what is buoyancy
a force exerted by a fluid that opposes an objects weight
what does archimedes principle state
the submerged weight of fish = difference between fish weight and the water is displaces
2 main strategies fish evolved to increase buoyancy
1 - reducing dense body parts
2 - using buoyancy aids (lipids/gas)
what is the specific gravity of seawater, fish muscle and bone
Seawater: 1.026
Fish muscle: 1.050
Bone: 2.000
what is dynamic lift
lift generated by movement (e.g. pectoral fins in sharks)
what is static lift
buoyancy provided by low-density substances e.g. lipids/gas
what is a major lipid used by elasmobranchs for buoyancy
Squalene (~0.86)
what law explains how gas volume changes with depth
Boyles Law
what are the two types of gas bladders
physostomous - bladder connected via pneumatic duct
physoclistous - duct closes; uses gas gland and rate mirabile
what is the function of the gas gland
secretes lactic acid, lowers pH and causes oxygen to dissociate from haemoglobin
what is the rate miracle
a counter current capillary system (helps inflate swim bladder)
What physiological effects help release O₂ into the bladder?
Bohr effect: Lower pH reduces Hb-O₂ affinity
Root effect: Reduces total O₂ carrying capacity
Salting out & CO₂ effects also reduce solubility and O₂ binding
why might deep-sea fish have multiple gas glands and long rete mirabile
to increase oxygen multiplication at high pressures for buoyancy control
why do some fish (scombroids) lose their swim bladder
to allow rapid vertical movement and avoid sonar detection by predators
what are two main steps in fish respiration
ventilation (moving water to gills) and extraction (taking oxygen from water)
what are the functions of gills
gas exchange
osmoregulation
acid-base balance
excretion of nitrogenous waste
structure of gill
gill arch
gill rakers
gill filaments with lamellae
what are two main types of ventilation
Ram ventilation (swimming forces water over gills)
Buccal ventilation (muscular pumping of water)
what adaptation do ram ventilation fish like tuna have
Gill fusions to improve efficiency
where do fish have rete mirabile
swim bladder
red muscle
retina
what is rete mirable
a counter current capillary system (helps to multiply gas/heat)
what is the role of the choroid gland
maintains high oxygen levels in the retina via the rete mirabile
what is a tapetum lucidum
a reflective layer behind the retina that enhances vision in low light
what are tubular eyes
fixed cylindrical eye adapted for upward, binocular vision (deep sea fish)
what is a neuromast
a sensory organ in the lateral line that detects water movement
what are otoliths used for
balance, hearing (aging fish)
what fish structure can amplify sound
swim bladder
what are ampullae of lorenzini
specialised electroreceptors in elasmobranchs and some teleosts used to detect electric currents
what are the two types of colour in fish
bio chromes - pigment based
schematochromes - structural, reflective
what hormones regulate physiological colour change
Melanin concentration hormone (MCH)
a-Melanocyte Stimulating Hormone (a-MSH)
what is disruptive colouration (silvering)
patterns like stripes/bars that break up the fish’s outline and confuse predators
what is facultative mimicry
ability to change colour/pattern to mimic multiple species
the two main sources of bioluminescence are …
symbiotic bacteria and self-luminous (photophores)
how do deep-sea fish use red light
produce and see red light to hunt prey like crustaceans
what is homeostasis
maintaining a stable environment including water and salt balance
what is osmoregulation
active regulation of water and ion concentrations to maintain internal balance
what is osmosis
movement of water across a semi permeable membrane due to solute concentration differences
what’s the difference between hyper osmotic and hyposmotic
hyper - fish is more concentrated than environment (water enters)
hypo - fish is less concentrated than environment (water leaves)
how do elasmobranchs osmoregulate
retain TMAO and urea to stay slightly hyper osmotic to seawater
Are marine teleosts hypo- or hyperosmotic to seawater?
Hyposmotic (must drink seawater)
how do freshwater fish osmoregulate
dont drink, produce dilute urine, β-chloride cells in gills to absorb salts
what are the three main types of jaw suspension in fish
amphistyly
autostyly/holostyly
hyostyly (sharks and bony fish)
what are pharyngeal teeth used for
grinding food
how is intestine length related to diet
carnivores - short intestine
herbivores - long intestine
what is hermaphroditism
having both male and female reproductive organs