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community
group of species that occur together in space and time
physically, taxonomically, statistically, interactively
how communities can be defined
richness
number of species
evenness
equability of individuals among species
diversity indices
combination of richness and evenness
rank abundance curves
shows relative species abundance in an ecological community
Shannon-Wierner Index
measured richness + evenness
Simpsons Index
measures dominance
predation
one group is negatively affected while the other is positively affected
mutualism
both groups benefit
amensalism
one group is negatively affected while the other group is neutral
commensalism
one group is positive affected while the other group is neutral
apparent competition
indirect interaction where two prey species negatively affect each other because they share a common predator, rather than directly competing for resources
trophic cascade
the top predator controls the structure of an ecosystem by negatively affecting prey and positively affecting producers
Grinnell’s niche
habitat or environment an organism is capable of occupying
Elton’s niche
the role a species plays in the community
competitive exclusion principle
two species cannot coexist on one resource
Hutchinson’s niche
“n-dimensional hypervolume” based on biotic and abiotic components
fundamental niche
the full potential range of environmental conditions and resources a species can occupy, only based on climate tolerances
realized niche
restricted range a species actually occupies, limited by competition and predation
niche partitioning
competing species divide resources so they can coexist instead of outcompeting each other
limiting similarity
there is a limit to how similar two species can be in their niches and still coexist
competition
the simultaneous demand by two or more individuals or species for an essential shared resource that is in limited supply. mutually negative interaction
intraspecific competition
competition between members of the same species forces them into a wider range
scramble competition
lack of a resource harms all organisms
contest competition (interference)
the best competitors get enough resources
interspecific competition
reduction in population growth rate of one species due to the presence of another species
allelopathic activity
toxins that reduce growth or seed production of other plants
asymmetric competition
uneven division of resources; large competitors have greater effects
consumptive competition
use of a shared, renewable resource
pre-emptive competition
sessile organisms acquiring space
overgrowth competition
one individual overgrows and blocks resources to another
chemical competition
use of toxins that diffuses over environment (allelopathy)
territorial competition
active defense of space
encounter competition
transient interactions over a resource
Lotka-Volterra Model of Competition
measures interspecific and intraspecific competition. (a>1= greater interspecific, a<1= greater intraspecific)
Zero Net Growth Isocline
values of populations 1 and 2 that yield zero population growth for both species
founder control
outcome depends on starting densities, more abundant species will dominate
resource
any substance that leads to increased growth rates as its availability is increased
law of the minimum
the resource with lowest supply limits growth
R*Theory
minimum equilibrium resource requirement needed to maintain consumer population growth
R*Rule
species with lowest R* should competitively exclude all other species
substitutive/replacement
best for comparing relative strengths of intra and interspecific competition
additive
best for measuring magnitude of interspecific competition
response surface design
best for comparing magnitudes of intra and interspecific competition
removal experiments
often easier to remove rather than add competitors
target-neighbor study
single individual of one species vs. natural densities of competitors
Hutchinsonian Ratios
coexisting species differ in size by a factor of 1:3
Paradox of the Plankton
found that aquatic habitats provide many more niches than originally thought, homogeneity rarely exists, and plankton communities may never reach equilibrium.
niche partitioning
natural selection drives competing species into different patterns of resource use or different niches
niche shifts
changes in a species diet or habitat use due to the presence of competitor (indirect evidence for competition)
character displacement
morphological differences between species that allows for coexistence
Townsend niche
a summary of an organisms tolerances and requirements