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natural selection
changes in phenoT → changing frequencies genoT
Q: best way model evol. outcomes
A: track the fitness of competing alleles
evolutionary stable strategy ESS
ANY B or char., once common in pop ≠ displaced by ANY competitor
UNlikely to be evolutionary stable
in free-living organims → innovation in group/ species @ expense individual
→ adaptions in nature = well-designed individual’s genes
selection CAN favour adap for higher lvl collections
→ mechanisms evolved abolish diff in reproductive success in constituent elements → + covariance between
collective’s functioning
reproductive success
covariance btw fitness of diff genes WITHIN organism x always total
→ intragenomic conflict & patterns that benefit genes @ expense of the whole organism