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Wavelength of visual light
400-700 mm
blue light has shorter wavelength, higher frequency
red light has longer wavelength, lower frequency
Particle properties of light
Tubes
One direction of light can be absorbed by each point
Lens
refracts/converges light
focal point
where an image will be reconstructed after being refracted through a lens
Pinhole optics
tiny hole only allows one point of light from each point on the actual object
image is projected upside down
smaller hole -> better resolution, lower brightness
Camera obscura
large pinhole camera in SF
Refraction
changing the angle of light by slowing it down
light moves at different speed through different substances
Refractive index
how much light bends towards a surface
proportional to the ratio of the speeds
power of lenses are measured in dioptecs = 1/f,
where f = focal length
more powerful lenses focus closer/have smaller focal length
Converging prism
a prism that converges light by refracting it inward
Photoreceptors
Rod cells and Cone cells
In the retina
Humans have 260 million receptors
Convert visible light into signals that can stimulate biological processes -> transduction
Rods
Rod shaped, smaller, used to see in the dark and perceive shapes, all one type, higher ration of rods to cones at the periphery
Cones
blue/green/red sensitive cones used to see colors
only cones at the fovea, less towards periphery
diurnal animals have more cones than nocturnal mammals
don't work well in low light situations
Compound eye
found in basic animals
early eye, 500 million years old
multiple lenses each with one photoreceptor, tubes
e.g. most insects, mantis shrimp
Simple eye
one lens focuses light on many photoreceptors
e.g. squids, humans, scallops (and most vertebrates?)
giant squid's eyes are like human eyes but bigger/better
Sensitive to transparent things, polarized light, broader parts of the EM spectrum
Limpet
100 receptors, no lens
Pinhole camera eye
Chambered Nautilus
Can let in a lot of light
Mirror eye
uses reflective material on the back of the eye to focus
very rare
e.g. brownsnout spookfish
Amoebe/paramecium
1 receptor, no lens
Copilia
2 lenses, 2 receptors
back lens scans over the image plane of front lens, used to focus
Extromission
the early idea that light shoots from our eyes
Scotopic vision
low light vision
rods only
only periphery sees
only in black and white
poor resolution
Photopic vision
bright light vision
only fovea/cones
good resolution
in color
Squid eyes
giant simple eyes, like human eyes but bigger/better
Sensitive to transparent things, polarized light, broader parts of the EM spectrum
Pupil
the pinhole, lets light in
Cornea
focuses light
lens
fine tunes focus
Iris
muscle that controls the size of the pupil
Retina
photoreceptors, where transduction from light to biological signals happens
Converging light
light rays that are parallel to each other are bent towards each other, focused
Optic function of an eye
form an image of an object on retina: cornea and lens work together to converge light to it
Optic nerve
Where nerves come together and exit the eye towards the brain
no photoreceptors where the optic nerve connects to the retina --> creates the blind spot
Fovea
the central focal point in the retina, around which the eye's cones cluster
periphery
optic disk
Area of the retina without rods or cones, where the optic nerve exits the back of the eye.
Types of cones/rods
1 type of rod
3 types of cones (correspond to R, G, B)
Are cones or rods larger?
cones
How many types of cones vs how many types of rods?
3 cones, 1 rod
Where are cones located?
mostly in the fovea
blood vessels in eye
cover retina, pushed away at fovea avascular zone (.5mm in diameter)
Adaptation
Our vision is most sensitive to change, no change in stimulation causes fatigue or adaptation in photoreceptors
This is why we don't see our blood vessels even though they are in front of the retina
Troxler fading
weak stimuli with no change--> brain ignores it
Receptive field of skin
area on skin that causes certain neurons to fire would be those neuron's RF
Photoreceptor RF
the area of the visual field that corresponds to single photoreceptor
receptive field of vision
area of your visual field that corresponds to an area on the retina
Neuron
a nerve cell
communicates using action potentials
can be very long
signals are very brief and fast
signals propagate to the end of the axon without decrement
what triggers an action potential?
a neuron will fire an action potential once the sum of it's IPSP's and EPSP's hit a certain threshhold
can be triggered with a strong input or lots of weak inputs
Action potential
the change in electrical potential associated with the passage of an impulse along the membrane of a nerve cell.
Neuron structure
soma (cell body) + dendrites -> axon -> presynamptic terminals-->axon terminal
what type of output does an action potential produce?
all or nothing
EPSP input or connection
Excitatory post synaptic potential
Makes neuron fire/Increases likelihood that neuron will fire
Depolarize a neuron
IPSP input or connection
Inhibitory post synaptic potential
Makes neuron not fire/Decreases likelihood cell with fire
Hyperpolarize a neuron
Depolarization
reduce the amount of negative charge in a neuron
Recording from neurons with an electrode
Sounds like rapid static
a neuron at rest has spontaneous activity
can measure depolarizations/hyperpolarizations to get a sense of when a neuron is firing
Excitation and inhibition counteract each other
summation of EPSPs and IPSPs trigger an action potential if cell hits the proper level of depolarization
Ganglion cells location
located past photoreceptors in retina
Ganglion cell RF
Ganglion cells have center/surround organization, on/off or off/on, measure local contrast
Preferred stimulus of an ON-center ganglion cell
bright light in middle of dark background, sensitive to local contrast
LGN RF
RF identical to ganlgion
Cells with orientation preference (how + which)
Simple cells, arrangement of LGN/ganglion cell RF's arranged with centers aligned
LGN location
thalamus
LGN input
take information from contralateral and a little from ipsilateral
LGN organization
organized topographically
LGN sends information to __ in the __ lobe
V1, occipital
Hermann grid RF explanation
Cells with RF that are near the corners of the grid signal low contrast because the junction of 4 paths impede the off surround
Why don't we see herman grid illusion at the center of our vision?
Fovea has smaller receptive fields
Simple cells: who discovered
Hubel and Wiesel, accident, with cat
Simple cell RF
on center/off surround, with bars not circles
Simple cell stimulus preference
Bars of light
Tuning curve for orientation
closer to orientation preference = higher response
Complex cell input
several simple cells
Complex cell preference
Bars of light that align with the RF, doesn't matter if it's dark or light, only sensitive to motion direction and orientation
Complex cell RF
Every location has an ON response from at least one simple cell and an OFF response from another simple cell
Preferences of complex RF's
position insensitive within the receptive field
contrast insensitive
orientation sensitive
direction of motion
Hypercomplex cells
same as complex cells but with length preference
end-stopped cells
have inhibitory regions surrounding RF
White matter
axons w/ myelin sheaths, show cells communicating with each other
Gray matter
cell bodies (soma)
Soma
cell bodies
Cerebral cortex
high level thinking and processing happens here (the lobes)
V1 location
Occipital lobe
Phineas gage
pole through frontal lobe
became an uninhibited, unreasonable jerk
Broca's area
Controls language expression - an area of the frontal lobe, usually in the left hemisphere, that directs the muscle movements involved in speech.
Phrenology
obsolete (and racist) theory that bumps on skull correspond to certain talents
Four lobes of the brain
frontal, parietal, temporal, occipital
Motor cortex
M1: frontal, precentral gyrus
Somatosensory cortex
S1: parietal
postcentral gyrus
Homunculus
Motor homunculus vs Sensory homunculus
size of body part is proportional to the proportion of the motor or sensory cortex devoted to it
area of cortex that corresponds to different body parts (either for motion or sensation)
Corpus Callosum
Located in the center of the brain: allows for communication between two hemispheres
white matter
Cortical magnification
parts of the body (hands, lips, face) are more represented on the cortex because more detailed sensory information
Tonotopy
areas in temporal
Somatotopy/ Motor topography
spatial mapping of the somatosensory and motor functions: adjacent areas correspond to adjacent body parts
retinotopy
mapping of visual input from retina to neurons, points that area adjacent on the retina are adjacent on cortex
Visual fields crossover
Contralateral organization of vision: 90% contralateral input, 10% ipsilateral
Walter Penfield
Neural surgeon who stimulated exposed brain of alert patients, used a probe to deliver slight electrical stimulation
different locations evoked different memories, sensations, or muscle twitches
mapped cortical homunculus
ventral pathway
What, occipital to inferior temporal lobe, object recognition
dorsal pathway
where, occipital to parietal lobe, location action naviagating and grasping
Medium scale modules
color, motion, facial recognition
note we can have deficits in all of these specifically b/c they are localized
Small scale models
Hypercolumns and columns
Hubel&Wiesel Hypercolumn structure
organized in slabs/ columns (orientation columns)
topographically organized
Hypercolumns
located in v1
each hypercolumns analyzes information from 1 region of the retina
fovea is overrepresented
one hypercolumn measures: light/dark, red/green, blue/yellow, size, orientation, motion, depth