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Darwin 2 main discoveries
1) all living things share common ancestry
2) species on earth evolve through natural processes
In order for natural selection to occur:
a. Populations are variable
b. Variants that are successful leave more offspring
c. Generation by generation species change
Evidence of Evolution
- fossil record
- classification and biogeography
- morphology & development
- DNA/
Fossil record
Demonstrates evolution through temporal succession
Faunal Succession
Relative age and correlation of strata
Fossil organisms follow one another in a definite and irreversible order. Simple forms proceeding to invertebrates then vertebrates
Linnaean Hierarchy
Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species ; nested groups... more closely related look most similar
Immortal genes
500; shared amongst all living species
Neoplatonism: view on fossils
Fossils produced in rocks by molding forces (Vis plastica)
Aristotelianism; view on fossils
Fossils are the product of spontaneous generation from non-living material
Middle Ages; view on fossils
Fossils regarded as works of the devil
Da vinci
Discovered that fossils start from organic matter & lie in positions that resemble living communities; fossils also to fragile to be carries great distances
Law of Superposition
oldest rocks at the bottom, youngest at the top
Stensen
Used tongue stones and real shark teeth to discover that they were not products of visplastica but actually remains of past sharks teeth
Paley
Eighteenth century English philosopher who proposed the idea of a conscious celestial designer of all life on earth
Evolution
Species undergo genetic change over time
Is evolution constant amongst species
NO: evolution does not predict constant evolution or speed at which it occurs
DEPENDS ON: evolutionary pressure each individual species experiences
Gradualism
Takes generations to produce a substantial evolutionary change
Strong natural selection
When evolutionary change can be fast
I.e.: animal or plant colonizes a new environment
Weak natural selection
Evolution often slows down
I.e.: When a species becomes well adapted to a stable habitat
Fundamental traits
- biochemical pathways we use to produce energy
- standard four-letter DNA code
- how code is read and translated into proteins
Speciation
Common ancestor x is the missing link between each species that evolved from it
Every pair of species shares a common ancestor sometime in the past; closely related species had a common ancestor that lived more recently
nested hierarchy
Big groups of species whose members share a few traits are subdivided into smaller groups of species whose members share a few traits are subdivided into smaller groups of species sharing more traits and so on down to species that share nearly all their traits
Diluvialism
Fossils as "proof" of biblical flood
Principle of original horizontality
Everything is layered horizontally
Principle of Lateral Continuity
layers are continuous until encountering an obstruction
Robert Hooke (1635-1703)
Suggested that fossils were remains of extinct organisms and that species have a limited lifespan
- fossils could be used to correlate strata
William Strata Smith
Published first series of high quality geologic maps of Britain correlating strata with the use of fossils
Faunal Succession
Fossils layer in rocks are older than the rock they rest in
Properties of good index fossils
- easy to identify
- geographically and environmentally widespread
- geologically short lived
Biological evolution
Organisms that change over time typically due to natural selection
Georges Cuvier & Alexandre brongniart
Generated maps for the Paris basin (1811) using faunal Succession
Erosional surfaces
Represent missing time & catastrophic events that happened earlier in earths history
Cuvier
Examined jaws of fossils found to modern day elephants and saw that they bone structure was the same
- believed life's moved progressively towards its "perfect" modern state
Catastrophism
The principle that events of the past occur suddenly and wildly
James Hutton
"No vestige of a beginning no prospect of an end"
Father of modern geology & uniformitarianism
-Used present processes to understand the past recorded in rocks (uniformitarianism)
-realized the temporal implications of the rock cycle (antiquity of earth)
Plutonism
Origin of igneous rocks is molten magma forcibly intruded upward into the earths crust due to subterranean heat
Enclosing relations
Intruding magma encloses older pre-existing rocks into its magma
Cross-cutting relations
Magma cuts through older existing rock
Principle of inclusion
Any rock represented by eroded fragments that are included in another rock must be older than the enclosing one
Uncomformity
Surface of erosion and or non-deposition separating two rock bodies; represents missing time
angular uncomformity
tilted rocks are overlain by flat-lying rocks
Nonconformity
metamorphic or igneous rocks in contact with sedimentary strata
Disconformity
A type of unconformity in which the beds above and below are parallel and sedimentary
Lyell uniformity of law
Natural laws are invariant in space and time
Lyell uniformity of process
Only processes operating today operated in the past
Wallace line
Between Asian islands and Australia
Placentals on one half & marsupials on the other half
Central tenents
Heritability
Variability
Differential reproduction
Excess
Divergence
Heritability
Offspring inherit a combination of traits from parents
Variability
Offspring not exact copies
Differential reproduction
Slight adaptive advantage greater chance of producing offspring
Excess
Many more offspring by everything that can possibly survive and be sustained
Divergence
Branching process
Hypothesis
A proposed explanation for an observed phenomenon...MUST be testable
Theory
A hypothesis that has withstood extensive testing by a variety of methods and in which a higher degree of certainty may be placed
Evolutionary predictions
1. Able to find some evidence for evolutionary change within the fossil record
2. Able to find some cases of speciation in the fossil record, with one line of descent dividing into two or more
3. Find examples of species that link together major groups suspected to have common ancestry
4. Species show genetic variation for many traits
5. Cases of imperfect adaptation
6. See natural selection acting in the wild
Darwin observations while on voyage
1) fossil animals are now extinct
2) distinction between species and varieties of species were sometimes unclear
3) geographic variation and species replacement
4) different varieties on different islands
5) South American affinities of Galapagos island populations
Stabilizing selection
Selects against extremes in a population (end members of populations)
- results in maintenance of the status quo around optimum
-can decrease variability in population
Directional selection
A type of natural selection where a single phenotype is favored, causing the allele frequency in the population to shift in one direction over time; One extreme selected against
TYPE OR INTENSITY OR DIRECTION OF SELECTION MAY CHANGE
Fitness
Relative ability to contribute offspring to next generation
disruptive selection
Natural selection where extreme forms are favored over intermediate forms
Selection for extremes in a population
- results in divergence away from existing mode
Drift
No selection present
- no differential reproduction, all forms equally successful
- expectation is no change in mean morphology of population
- typically results in an increase in the variability of a population
Mutations
Copying errors that randomly occur during DNA replication as cells grow and divide
Natural selection and randomness
Natural selection involves the nonrandom survival of randomly generated variants
Bacteria
Do not reproduce sexually; evolve quickly due to;
Cell-cell transfer genes
Large population sizes
Rapid generation times
Biological species concept
Group of inter-breeding natural populations that are reproductively isolated from other such groups
Artificial selection
The process by which a species is modified via human actions that encourage the breeding of individuals with certain traits
Human intervention
Controlled, selective breeding of domestic animals
Why did Darwin need artificial selection as an analogy to natural selection
Because natural selection is often too slow for us to observe in a lifetime
Because artificial selection has resulted in dramatic changes
Because artificial selection demonstrates inheritance
Gregor Mendel
Father of genetics
Raised > 29000 pea plants between 1856-1863 &a closely controlled reproduction and kept careful notes on plant traits generation by generation
Bending inheritance
Each hereditary factor is permanently diluted in a hybrid (WRONG!!!)
Mitosis
Normal cell division
Meiosis
Cell division that produces reproductive cells in sexually reproducing organisms
Locus
A given site on the chromosome
Alleles
Alternative forms of the same gene
Homozygous
2 alleles are identical
Heterozygous
2 different alleles
dominant allele
Is expressed even when heterozygous
Recessive allele
Is only expressed when homozygous
Mendelian Genetics
- phenotype traits are determined by units of heredity called genes
- each parent contributes one randomly chosen allele of each gene to each offspring (principle of segregation)
Wrong aspects of Mendelian genetics
-One gene codes for one trait
-Some alleles are dominant others are recessive
-Each pair of alleles (gene) behaves independently of other allele pairs (principle of independent assortment)
incomplete dominance of alleles; what is it and what is it an example of?
A condition in which neither allele for a gene completely conceals the presence of the other; results in intermediate expression of trait I.e flower color
EXAMPLE OF NONMENDELIAN CHARACTERISTICS
Con-dominance of alleles; what is it and what is it an example of?
Both alleles fully expressed i.e AB blood type
EXAMPLE OF NONMENDELIAN CHARACTERISTICS
Many alleles for one gene ( in a population)
Non Mendelian characteristic
One gene may have multiple effects
I.e allele for albinism
Non Mendelian characteristic
One trait may result from the input of many genes
I.e cow milk production & human skin color
Non Mendelian characteristic
Genes may influence each other
I.e hello lab fur
Non Mendelian characteristic
DNA
Deoxyribonucleic Acid
4 basic bio molecules of life
1. Carbohydrates (CHO): fast energy
2. Lipids (CHO- glycerol, fatty acids): long term energy storage, cell membranes
3. Proteins (CHON, made of amino acids): builds tissues, muscles, immune systems and enzymes
4. Nucleus acids (CHONP): DNA, RNA
DNA structure
Ladder twisted into a double helix
- holds information to build proteins and direct development
DNA sides
Sugar-phosphate backbone
DNA rungs
Base pairs
Base pairing rules
Adenine pairs with Thymine
Gaunine pairs with Cytosine
Nucleotides
Basic units (building blocks) of DNA molecule, composed of a five carbon sugar, a phosphate group, and one of 4 DNA (nitrogenous) bases
Anti parallel strands
DNA strands run in opposite directions
Why study DNA
- is the fundamental basis of variation and change
- DNA operates the same way in ALL living things
- many current issues involve an understanding of DNA
DNA replication process:
- Begins by unzipping the molecule
- a complimentary base hooks up to each unpaired base
- result is two daughter DNA molecules; each is half old half new
Helicase
An enzyme that untwists the double helix of DNA at the replication forks.
DNA polymerase
Enzyme involved in DNA replication that joins individual nucleotides to produce a DNA molecule
Gene
Segment of DNA molecule that codes for an amino acid chain to build a protein
Where does DNA replication occur:
Inside the nuclear and before an individual cell divides