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Assemblage
a subset of a community, a group of species, usually of the same taxa, done at the assemblage level becuase studying a whole community is difficult.
Forbes- 1887
“the lake as a microcosm” proposed lakes as a coherent unit. “the balance of nature” Organisms work together to balance the system (population size)
Clements 1916
Agreed with Forbes, plant succession and ecosystem changes. Proposed communities as coherent units. “super organism concept” communities are born, grow, and die like organisms. Directions among species keep species in certain areas, mainly focused on biotic interactions like competition.
Gleason 1926
Individualistic concept- each species has their own requirements and tolerances to survive/ grow. If species occur in the same area together it is because they respond the same way to the environmental gradient (overlap). Species abundance vs env gradient (no transition zones). He focused on abiotic factors.
Elton 1927
Animal ecologist, focused on predator/ prey interactions (food web/ invasion ecology) Proposed that communities have limited membership.
Lotka 1925 and Voltara 1926
Developed equations to describe interactions mainly competition. Showed that when two species compete for the same limited resource they cant co exist.
Gause 1934
Tested Lotka and Voltara models on 2 species of paramecium. Developed the competitive exclusion principle. Hard to apply to diverse communities
Grinnell 1917
Niche is the habitat or ecosystem that an organism can occupy.
Elton 1927
Niche is the role an organism plays in a community, focused on trophic niche.
Gause 1943
Made connections between degree of niche overlap and intensity of competition.
Hutchinson 1957
Proposed a way to quantify niche (n-dimensional hypervolume). Distinguished between fundamental and realized niche
Velland 2010
Conceptual framework of the functioning of community ecology, the black box of community ecology
4 Processes of distribution and abundance
Selection, speciation, drift, and dispersal.
Selection
anything that determines relative success of a species in a community. A determinate ( non random) process result of competition, disease, predation, etc…
Speciation
Evolutionary process creates diversity/ new species
Drift
Stochastic (random) changes in relative abundances, due to chance, leads to random chance of success, common in birds/ bats.
Dispersal
Does not consider migration, refers to movement in or out of local habitats for things like less competition, or more resources. Leads to gene flow/ species persistance.
Whittaker 1960/1972
Proposed diversity can be measured on different spatial scales like alpha, beta, and gamma diversity.
Alpha diversity
represents number of species in a local community/ smaller size area within a habitat.
Beta diversity
A measure of the different species composition between 2 or more habitats within a region.
Gamma diversity
total number of species in a region.
LDG time theory Wallace 1878
The tropics are older and have been habitable for longer since they were not effected by glaciation, they have had more time to accumulate species.
LDG Productivity theory
There is higher energy from the sun which increases primary production, thus increasing total ecosystem productivity and feeding higher trophic levels more effeciently.
Colwell and Hartt 1994
Proposed LDG reflects the outcome of putting species in a bounded domain. Pencil box model.
LDG More individuals hypothesis
The higher primary productivity leads to higher species richness which allows for increased speciation.
LDG tropical niche conservationism theory
Lineages originate in the tropic and stay in the tropics, species lack the flexibility to evolve and adapt to temperate climates.
LDG Evolutionary hypothesis
main focus is on diversification (speciation- extinction) rates
Stebbins 1974
Proposed the tropics are a cradle for the generations of new species and a museum for the preservation of the taxa there. High potential for speciation and low extinction rates= high diversification rates.
Mechanisms of productivity
selected species, niche complementarity, and competition.
Niche complementarity
resource partitioning by different species increases productivity through facilitation and resource partitioning
Nutrient cycling and retention
Niche complementarity increases resource partitioning, which increases SR. This increases resource use and efficiency which decreases nutrient retention.
Exponential model assumptions
remove I/E, closed population
population growth is constant, time steps are very small
constant b/d rates
no genetic structure
no age structure
Logistic model assumptions
b/d are linear
closed population
no genetic/ age structure
continuous growth
constant k
Log growth model ( small oscillations)
rT is less than 0.37
Medium product (dampened oscillations)
0.37< rT< 1.57
Large stable cycle
rT>1.57