the nerve cord is surrounded by a series of hard vertebrae made of either cartilage or bone
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Infraphylum Hyperotreti
retain notochord through adult stage; lack jaws, mouth with 4 paired fins; possess slime glands and are excessively slimy
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Class Petromyzodontida: Lampreys
ancerstors are ostracoderms, first known vertebrates, lack jaws, notochord in juveniles only, no paired fins, have skeleton of cartilage
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class Chondrichthyes: cartilaginous fishes
ancestors are placoderms, first known jawed vertebrates; Chondrichthyes appear around 370, key innovation: jaws; paired fins, have 2 chambered heart, have skeleton of cartilage, skin covered by placoid scales
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class Osteichthyes: bony fishes
ancestors are placoderms, first known jawed vertebrates, largest group of vertebrates; key innovation: Skeleton made of bone; possess jaws, paired fins, have 2 chambered heart, skin covered by dermal scales, 2 groups: ray-finned fishes and lobe-finned fishes
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Class Amphibia
ancestors are lobe-finned fishes; primarily terrestrial as adults and aquatic as juveniles, key innovation: possess 4 limbs, have 3 chambered heart, skin lacks covering; may be used for breathing, lack an amniotic egg
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Class repitilia
first appear around 315 my ago, primarily terrestrial but some aquatic, key innovation: amniotic egg; have 3 or 4 chambered heart, skin covered by scales
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anapsids
have no openings or fenestra behind the eye socket
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synapsids
have a single fenestra
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diapsids
have two fenestrae
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class reptile: birds
formerly class aves, first appear 150 my ago, closely related to one group of dinosaurs, key innovation: body covered with feathers, endothermic, amniotic eggs, possess wings
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bird ancestry
archaeopteryx lithographica
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avian characteristics of birds
feathers, endothermic, claws on feet, hollow bones, oviparous, sexual dimorphism
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Why are birds in class reptilia?
heterogametic females, feathers are modified scales, primitive birds had teeth, scales on feet and legs, lay eggs, claws not nails, nictitating membranes
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class mammalia
first appear around 160-200 my ago, evolved from a group of early reptiles, key innovations: hair, mammary glands for feeding young, endothermic
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monotremes
egg-laying mammals
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marsupials
pouched mammals
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placental mammals examples
cows, mole rats, capybara, river dolphin
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definition of ecology by Haeckel
oikos (greek)- house; the study of interactions between organisms and the environment, defined by Haeckel
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Krebs definition of ecology
the study of interactions that determine the distribution and abundance of organisms
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Townsend, Harper and Begon definition of ecology
the scientific study of the distribution and abundance of organisms and the interactions that determine distributions and abundances
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distribution
where organisms occur and why
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abundance
how many organisms there are in a particular location
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interactions
how and why does an organisms interactions with other organisms and with its environment affects its distribution and abundance
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studying ecology of individuals
how do they adapt to or deal with changes in their environment
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studying ecology of populations
ecology of groups of organisms belonging to the same species
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studying ecology of communities
ecology of populations belonging to different species
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studying ecology of ecosystems
ecology of communities plus environmental effects
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studying ecology of ecosphere
interactions among living and non-living factors across the whole earth
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two ways to describe a populationâs distribution
1. extent of the habitat over which an organism occurs 2. where individuals occur within a population
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kinds of biomes
aquatic, desert, forests, grasslands, tundra
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three common distribution patterns among individuals within a population
random, regular, clumped
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random distribution
each individual is equally likely to occur anywhere, there are no strong effects of competition or the environment on the distribution
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regular/uniform distribution
each individual is uniformly spaced within the population, this can be due to competition among individuals for resources or to some regularity in the distribution of resources
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clumped distribution
individuals occur in high density patches, separated by areas with few individuals; can be due to mutual attraction among individuals or attraction to a common resource
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measuring abundance
total number of individuals, density = number of individuals divided by unit area (volume)
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typically we ________ abundance rather than count every individual becauseâŚ.
estimate; too hard to catch or find all individuals, too time-consuming to count all, too expensive to count all
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space based methods
divide the population into equal sized areas/volumes, used for less mobile organisms
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mark recapture methods
used for mobile organisms
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why do we try to estimate population sizes
we would like to know how populations change in size overtime
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problems with exponential growth
population increases forever
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logistic growth equation
N=rN (1-N/K)
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equation for population size
Nfuture= Nnow + B-D+I-E
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Nnow
the population size now
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Future
the population size in the future
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B =
there number of births between now and the future
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D =
the number of deaths between now and the future
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I =
the number of immigrants. between now and the future (individuals entering population)
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E=
the number of emigrants between now and the future (individuals leaving the population)
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change in population size (from time now to time in future) is equal to the
difference between the number of births and the number of deaths
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B>D
population size goes up
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B
population size goes down
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birth and death rates equation per individual
ďN = (b âd)N
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what does r stand for in rN
the intrinsic rate of population increase or population growth rate
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if r>0
population grows
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if r
population shrinks
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K=
carrying capacity, maximum population size a habitat can hold
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growth is nearly exponential at
low population size
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growth is very slow at
high population size
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the logistic equation is more
realistic and suggests that populations cannot grow indefinitely but will instead reach some maximum size
long life, slower growth, late maturity, fewer large offspring, high parental care and protection, high investment in individual offspring
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life table
look at population growth in some more detail, divides the population into distinct groups (ages or stages)
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what is the most common life table
cohort table - follow all offspring born at a given time from birth until the last individual
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what information can you get from a life table
population age structure
population growth rate
population survivorship patterns
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life table variables
x
nx
lx
bx
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x=
the age or stage
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nx=
the number of individuals in each age/stage
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lx=
the percent of the original cohort that survives to age/stage x
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bx=
the number of offspring produced per individual in age/stage x
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If we add up all the values of in the lxbx column we can obtain the
net productive rate or R
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survivorship curve
a plot of the x values on the horizontal axis and the lx values on the vertical axis in a life table
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Type I survivorship curve
occurs when the most mortality occurs late in life
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Type II curve
occurs when mortality is spread equally across different stages
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Type III curve
occurs when most mortality occurs very early in life
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the demographic transition hypothesis
human societies tend to follow a pattern in population growth as they become more advanced
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stage 1 of the demographic transition hypothesis
high but nearly equal birth and death rates, r is low
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stage 2 of the demographic transition hypothesis
death rates decline but birth rates stay high, since b>d population increases
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stage 3 of the demographic transition hypothesis
death rates decline as well but b is still > d
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stage 4 of the demographic transition hypothesis
low but nearly equal birth and death rates, so r is low
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fully domesticated mammals have undergone parallel changes
increased color variability
pedomorphosis (retain juvenile traits)
brain reduction
docility
earlier maturation
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Who proposed that there were stages in the domestication of a species
Zeuner (1963)
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Stages in the domestication of a species
loose ties, captivity, intentional breeding, planned development, elimination of the wild species as competitors
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habitat
the physical area in which an organism lives
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niche
a description of all factors which influence an organismsâs growth, survival, and reproduction
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fundamental niche
any set of conditions under which an organism can occur and survive
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realized niche
the conditions under which an organism really does occur
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the fundamental niche is always
as large or large than the realized niche
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neutralism
effect on x= 0
effect on y= 0
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amensalism
effect on x= 0
effect on y= -
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commensalism
effect on x= +
effect on y=0
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competition
effect on x=-
effect on y=-
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mutualism
effect on x= +
effect on y=+
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predation or parasitism
effect on x= +
effect on y=-
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definition of competition
an interaction between individuals for a shared resource in limited supply which leads to a reduction in survivorship, growth, or reproduction for at least one of the individuals, cost for both
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contest competition
individuals fight for access to a resource, may involve direct contact or just threats
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scramble competition
individuals do not directly interact, try to get resources more quickly than others do