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3 conceptual models of succession
1. facilitation (primary succession)
2. inhibition (secondary succession)
3. tolerance (secondary succession)
Facilitation
1. Barren ground uninhabitable by all but most stress tolerant colonists
2. Early colonists make environment more suitable for successive species
3. Sequence continues until most competitively dominant species no longer facilitate invasion and growth of other species
inhibition
1. Initial community composition is who gets there first
2. Once colonist becomes established, it inhibits growth of following arrivals
3. Only when space and/or resources released through decay or decay of dominant residents can new colonists invade and grow
Tolerance
1. initial community composition is who gets there first
2. specis that appear later arrived later, or arrived early but grew more slowly
3. late arriving species tolerate presence of early species
- good competitors
- over time, late successional species exclude others
4. early successional species have no effect on late successional species (can live together)
Allogenic succession
When primary forces driving succession are from outside the system.
- immigration of new species
- seasonal changes in weather/sunlight
- disturbance
Autogenic succession
A type of succession where the plant community changes the environment; changes that occur within a community caused by the actions of the community members
- biotic
-has to occur within a forest
Primary succession
Starts on surface with no soil and usually no prior life.
ex. bare rock, lava rock
Secondary succession
Succession after non-catastrophic event, disturbance but soil still there.
Ex. after forest fire, flood
Intermediate disturbance hypothesis
Species diversity is highest at intermediate levels of disturbance because both early and late-successional species can coexist.
Behavioral cascade
Predator alters foraging behavior of its prey which stimulates primary production or alters population size lower trophic levels.
Ecosystem engineer
Species that physically modifies habitat and changes resource availability for other species.
Allogenic engineer
Changes environment by transforming materials from one state to another (e.g., beavers building dams).
Autogenic engineer
Changes environment through own physical structure (e.g., corals).
Trophic cascade
Top down control where higher trophic levels regulate abundance of lower trophic levels, control of primary production is biotic (consumers)
Bottom-up control
Resource availability at base of food web controls abundance of higher trophic levels, control of primary production is environmental abiotic
Trophic energy transfer
Only about 10% energy transferred from one trophic level to the next, so 90% is lost.
Headwater streams/ lakes
Fueled by allochthonous inputs (outside) and useful for testing bottom up effects bc boundaries and nutrient imputs easier to measure
Productivity hypothesis
argues that more productive ecosystems should have longer food chains/bottom up contorl determines food chain length
Ecosystem size hypothesis
Food chain length increases with ecosystem size
Productive-space hypothesis
Food chain length increases with ecosystem productivity and size
Exploitation ecosystem hypothesis
food chain length is controlled by bottom up forces, with top down control becoming increasingly important as more trophic levels are added
ex. herbivory, parasitism, predation
3 R's of Community stability
Resistance, return time, resilience (persistence).
Resistance
How much the community changes due to specific disturbance.
-more resistance=more stable
-How big a change did the disturbance cause?
Return time
Time it takes a community to return to equilibrium state.
- how quickly did the community recover from disturbance
Resilience (persistence)
How closely the post-recovery community resembles the pre-disturbance community.
Diversity vs. stability
More habitat and species diversity generally increase stability and persistence after disturbance.
Food web theory
Communities with higher connectance (with many species interactions) should be more stable.
How does biodiversity loss affect community stability?
More omnivores you have, more stable the environment/food web
Connectance
Number of realized interactions among species in the food web.
Keystone species
Effect on community structure is much larger than expected from its abundance or biomass.
Invasive species
Non-native species that spreads and causes ecological, economic, or human health harm; alter community structure
Succession
The series of predictable changes that occur in a community over time
r-selected species
-many offspring
-shorter life
-early reproduction
-pioneer species
K-selected
-density dependant
-fewer offspring
-longer life
-climax community
Climax community
stable, mature ecological community with little change in the composition of species
Disturbance Regime
timing, magnitude (small or large), frequency, predictability
Alternative state
occur when more than one type of community can exist in a particular environment