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What Direction is Primary Growth?
Up
Apical Meristem
From the Tips- Roots and Shoots. Produces primary tissues
Lateral Meristem
Side
Vascular Cambium
Growth in Diameter
Cork Cambium
Growth in Bark
Secondary Growth
Outward
Primary Tissues
Epidermis
Cortex
Primary Phloem
Primary Xylem
Pith
Where does Vascular Cambium Grow
Between Xylem and Phloem
What are the two types of cambium initials
Fusiform initial
Ray initial
Ray initial
beginning of rays. Horizontal growth
Fusiform inital
Vertical Growth
Perclinal division
division that splits cambium cells into one cambium and one xylem
increases diameter
Anticlinal division
division that splits cambium cells into two cambium
increases circumference
Wood Cell Growth Process
Cambium
Enlarge
Secondary Wall
Ligninification
Death
How many rings of Juvenile Wood
8-25 rings
What factors influence the amount of Juvenile wood
Species
Genotype
Age
Environment
Growth Conditions
Juvenile Wood Specific Gravity
LOWER specific gravity
Juvenile Wood Longitudinal Shrinkage
HIGHER Longitudinal Shrinkage
Juvenile Wood Radial Shrinkage
LOWER radial shrinkage
Juvenile Wood Density
LOW density
Juvenile Wood Strenth
WEAK
Juvenile Wood Chemical Properties
High Moisture
Higher Lignin and hemicellulose
Low cellulose
Juvenile Softwood
Short thin trachieds
Low Density
Larger MFA
High lignin, low cellulose
Juvenile Hardwood
Short thins FIBERS
Low Density
High Holocellulose
How is Heartwood Formed
When Sapwood Stops Conducting Water
What can cause Sapwood Blockages
Air Bubbles
Pit Asperation
Tyloses
Extractives
What factors control Parenchyma death
Spatial Control
Temporal Control
Spatial Control
How far from cambium
Temporal Control
Age of the Cell
Extractives
Organic compounds made by parenchyma cells
Effects Wood Permability+Toxicity
Good Effects of Extractives
Good Color
Durability
High Compressive Stability
Bad Effects of Extractives
Hard to Pulp
Tannins corrode metal
Colors mitigate while drying
Interferes with paint, glue, etc drying
Cell Wall Element %
Carbon 49%
Hydrogen 6%
Oxygen 44%
Nitrogen >0.1%
Ash <0.1%
Cell Wall Organic Polymers %
Cellulose 50%
Hemicellulose 25%
Lignin 25%
Cellulose
Most Abundant Polymer on Earth
Long Straight Chain 10um
Strong Along Axis
Hemicellulose
Branched Chain Polymer
Variable in composition
Made of Sugar Units
Lignin
Made of Phenylpropane
Stable but difficult to isolate
Resistant to decay
Provides Structure
What are Microfibrals
Cellulose fiber structures
Cellulose in 1 Fibral
40-100 Cellulose
Fibrals in a Microfibral
4-6 Fibral
Compound Middle Lamella
Intercellular Substance made of primary walls and true middle Lamella
Pectin + Lignin
.2-2 um
Primary Wall
Thin and Stretchable
Microfibrals randomly arranged
Hemicellulose, Pectin, Lignin
Secondary Wall
Made of 3 Layers- S1,S2,S3
S2 largest layer 5-10um
Cellulose, Hemicellulose, Lignin
Size of Secondary Wall Layers
S1: 4-6 Lamella
S2: 30-150 Lamella
S3: 0-6 Lamella
Tertiary Wall
Not always present
Warty Layer- Softwoods
Gelatinous Layer- Hardwoods
Microfibral Angle (MFA)
Primary: Random
S1: 50-70 degrees
S2: 10-20 degrees
S3: 60-90 degrees
Reaction Wood
Forms as a Reaction to an event
Returns Stem to original angles
Compression Wood
Forms in Softwoods
Lower side of the tree
Higher Lignin 8-9%
Lower Cellulose 10%
Cellular Properties of Compression Wood
Rounded Trachieds
EW Trachieds have thicker cell walls
Short Trachieds with smaller pits
no S3
Physical Properties of Compression Wood
Higher Density 10-40%
High Compression Strength
Lower tensile and impact bending
Warps while drying
Tension Wood
Hardwoods
Wider Growth Rings
Adds G-Layer
G-Layer
Gelatinous Layer
Forms inside or replaces S3
Tension Wood Chemical Properties
Less Lignin 8-9%
More Cellulose 5-10%
Tension Wood Physical Properties
Higher Density 5-10%
Higher Longitudinal Shrinkage 1-2%
Collapses while Drying