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What are the three major building blocks required to make plant tissue?
Carbon (CO2)
Which elements make up most of plant biomass?
Carbon
Where do plants obtain carbon?
From atmospheric CO2
Where do plants obtain mineral nutrients?
From soil
What are mineral nutrients?
Essential elements obtained from soil
Why are mineral nutrients also found in animals?
Plants are the base of food webs
How are plant nutrients classified?
Macronutrients and micronutrients
What distinguishes macro- from micronutrients?
Amount required
Why is nitrogen a macronutrient?
Required in very large quantities
Which biomolecules contain nitrogen?
Proteins
Why are fertilizers labeled NPK?
Nitrogen
What is Liebig’s Law of the Minimum?
Growth is limited by the scarcest resource
What analogy explains Liebig’s Law?
Shortest plank in a barrel
Which nutrients most commonly limit growth?
Nitrogen and phosphorus
What is global nutrient limitation?
Growth below climatic potential due to nutrients
Why are many old soils nutrient-poor?
Leaching and lack of renewal
Why are Australian soils infertile?
Very old geology
How do clay soils affect nutrients?
Strong binding
How do sandy soils affect nutrients?
Poor retention
What is nutrient homeostasis?
Regulated balance of uptake and use
What is chelation?
Binding toxic elements to reduce reactivity
What are heavy metals?
Non-essential toxic elements
Give examples of heavy metals
Mercury
Why are heavy metals toxic?
Enzyme inhibition and ROS generation
How can heavy metals enter plants?
Mimic nutrients
What determines metal availability?
Soil pH
Why does liming reduce toxicity?
Raises pH
What is symbiosis?
Close association between organisms
Does symbiosis imply benefit?
No
What symbioses affect plant nutrition?
Nitrogen fixation and mycorrhizae
What do plants provide symbionts?
Carbon and habitat
What do symbionts provide plants?
Nutrients and water
How old are mycorrhizal symbioses?
~400 million years
Why were mycorrhizae important for land plants?
Improved nutrient uptake
What nutrient limits most terrestrial ecosystems?
Nitrogen
Why is nitrogen abundant but limiting?
Exists as inert N2 gas
What bond makes N2 inert?
Triple bond
Which organisms can fix nitrogen?
Some prokaryotes
Can plants fix nitrogen directly?
No
What enzyme fixes nitrogen?
Nitrogenase
How much ATP does fixation require?
~16 ATP per N2
Why is nitrogenase anaerobic?
Oxygen inactivates it
Which plant family contains most nitrogen fixers?
Fabaceae
What are legumes?
Members of Fabaceae
Give examples of legumes
Peas
What bacteria fix nitrogen in legumes?
Rhizobia
Where are rhizobia housed?
Root nodules
Do rhizobia fix nitrogen freely?
No
What is traded in legume symbiosis?
Carbon for nitrogen
Why is signaling specific?
Avoid pathogens
What plant signal attracts rhizobia?
Flavonoids
What bacterial signal triggers nodulation?
Nod factors
What plant structure curls during infection?
Root hair
What is an infection thread?
Guided bacterial tunnel
What are bacteroids?
Differentiated rhizobia in nodules
What is facultative fixation?
Plants can regulate fixation
What are costs of fixation?
Carbon
Why is oxygen a problem?
Inhibits nitrogenase
What molecule manages oxygen?
Leghemoglobin
Why are nodules pink?
Leghemoglobin
How do plants sanction cheating rhizobia?
Smaller
When is fixation favored?
Low N
Why doesn’t fixation dominate ecosystems?
High cost and shared benefits
What is autoregulation of nodulation?
Systemic suppression of nodules
When is fixation highest in succession?
Rapid growth phase
What are mycorrhizal fungi?
Root-associated fungi
What do mycorrhizae increase?
Effective root surface area
Why are hyphae effective?
Thin and extensive
Which nutrient commonly moves via fungi?
Phosphorus
What are ectomycorrhizae?
External root fungi
Do ectomycorrhizae enter cells?
No
What are endomycorrhizae?
Intracellular fungi
What structures increase exchange?
Arbuscules
Which plants are ectomycorrhizal?
Conifers
Which plants are endomycorrhizal?
Most flowering plants
What special ability do ectomycorrhizae have?
Break down organic matter
How do mycorrhizae increase P availability?
Organic acids
How do mycorrhizae affect drought tolerance?
Increase water uptake
What hormone attracts mycorrhizae?
Strigolactones
What fungal signal confirms compatibility?
Myc factors
Why are mycorrhizae reduced in agriculture?
Tillage and fertilizers
What are biofertilizers?
Microbial inoculants
Why is regulation important?
Products vary in quality
What traditional system uses fixation?
Three Sisters
Which crop fixes nitrogen there?
Beans
What are endophytes?
Microbes living inside plants
Where were leaf N-fixers found?
Sugarcane
How was leaf fixation shown?
Radioactive N2 uptake
What is the rhizosphere?
Root-influenced soil
How do plants shape microbes?
Exudates and pH
What does PCA show?
Distinct communities
What are plant growth-promoting bacteria?
Beneficial rhizosphere microbes
What is a common mycorrhizal network?
Shared fungal connections
Is resource sharing proven?
Evidence inconclusive
Why are nutrient trade-offs important?
Limited resources
Why are nutrients key for climate models?
Limit carbon uptake